唇边书带蕨Vittaria elongata Sw.
唇边书带蕨Vittaria elongata Sw.
12. 唇边书带蕨（中国高等植物图鉴）长叶书带蕨（中国主要植物图说蕨类植物门）、阔叶书带蕨（中国蕨类植物图谱）、垂叶书带蕨（台湾植物志）、双唇书带蕨（西藏植物志）图版7：1-4.
Vittaria elongata Sw., Syn. Fil. 109, 302. 1806; Hook. et Bak. Syn. Fil. 395. 1874, pro parte; Bedd., Ferns South India t. 21. 1863 et Handb. Ferns Brit. India 404, f. 238. 1883; Clarke in Trans. Linn. Soc. Lond. II. Bot. 1: 573. 1880; Ching in Sinensia 1 (12): 179. 1931; H. Ito in Journ. Jap. Bot. 12: 466. 1936; Tard. -Blot et C. Chr. in Lecomte, Fl. Indo-Chine 7 (2): 197. 1940; Holtt., Rev. Fl. Mat. 2: 614, f. 360. 1955 et in Dansk Bot. Ark. 20: 34. 1961; 傅书遐, 中国主要植物图说 蕨类植物门 252, 图338. 1957; Bir in Res. Bull. (N. S.) Panjab Univ. 13 (1-2): 17, f. 1-5. 1962; Ching et al. in W. Y. Chun et al., Fl. Hainan. 1: 201. 1964; Ic. Corm. Sin. 1: 281, f. 561. 1972; DeVol in H. L. Li et al., Fl. Taiwan 1: 238. 1975; Y. L. Chang et al., Sporae Pterid. Sin. 375, t. 87, f. 5-6, 1976; Dixit in Journ. Econ. Tax. Bot. 2: 210, f. 1-9. 1981; Fl. Fujian. 1: 260, f. 246. 1982; Tagawa et K. Iwats., Fl. Thail. 3 (2): 223. 1985. ——Pteris graminifolia Roxb. in Calc. Journ. 4: 502, t. 33. 1844. ——Vittaria anodontolepis Fee, Mem. Foug. 3: 23, t. 4, f. 3. 1851-52. ——Vittaria boninensis Christ in Warb. Monsunia 1: 57. 1900. ——Vittaria ogasawarensis Kodama in Matsum., Ic. Pl. Koishikav. 3: 146. 1915; H. Ito, l. c. 467. 1936. ——Vittaria tortifrons Hayata, Ic. Pl. Forms. 6: 162. 1916; H. Ito, l. c. 466. 1936. ——Vittaria formosana Nakai in Bot. Mag. Tokyo 39: 176. 1925. ——Vittaria pauciareolata Ching in Sinensia 1 (1): 11. 1929 et in Hu et Ching, Ic. Fil. Sin. 1: pl. 38. 1930. ——Vittaria zosterifolia auct. non Willd. : Ching et al. in Chun et al., Fl. Hainan. 1: 202. 1964; Ic. Corm. Sin. 1: 281. 1972; DeVol in H. L. Li et al., Fl. Taiwan 1: 238. 1975; Y. L. Chang et al., Sporae Pterid. Sin. 377, t. 87, f. 7-8. 1976; Nakaike, Enum. Pterid. Jap. Fil. 84. 1975 et New Fl. Jap. Pterid. 716, cum f. 1992; Dixit, 1. c. 212, f. 8. 1981; Fl. Fujian. 1: 260. 1982; Ching et S. K. Wu in C. Y. Wu, Fl. Xizang. 1: 105, f. 26: 1-2, excl. 3. 1983; X. Cheng in W. T. Wang, Vasc. Pl. Hengduan Mount. 1: 64. 1993. ——Vittaria merrillii auct. non Christ: Ching, l. c. 180. 1931; Dixit, l. c. f. 9. 1981.
根状茎长而横走，多分叉，密被须根和鳞片；须根的根毛绒毛状，密布，形成线状的吸水结构；鳞片黑褐色，具亮光泽，钻状披针形，长4-5毫米，基部最宽，约0.5-1毫米，先端渐尖，纤毛状，端部呈腺体状，粗筛孔网眼壁增厚，黑色，边缘具较长的睫毛状齿；叶稍远生，相距约0.5毫米，通常成丛倒垂。叶柄横切面椭圆形，或长或短，基部常被较根状茎鳞片狭长的鳞片；叶片线形或带状，长可达1米以上，宽0.5-2厘米，顶端圆头或钝头，下部渐狭，全缘；中肋细，两面扁平，不甚明显，侧脉多数，形成斜升的网眼，较明显。叶近革质，干后有皱纹。孢子囊群线着生于叶缘的双唇状夹缝中，开口向外，自叶片的近基部延伸直达近顶端；隔丝多数，顶端杯状，长略大于宽。孢子狭长椭圆型，单裂缝，表面纹饰模糊。
分布于福建（永泰、宁德）、台湾（全岛分布）、海南（全岛分布）、广东（信宜、肇庆、阳春、阳江）、广西（南宁、龙州）、云南（河口、屏边、金平、勐腊、勐海、景洪、福贡、贡山）、西藏（墨脱）。附生于树上或林中岩石上，常同王冠蕨 Pseudodrynaria coronans (Wall. ex Mett.) Ching，鸟巢蕨属 Neottopteris 及铁角蕨属 Asplenium 等附生蕨类植物混生，海拔190-1350米。也分布于越南、泰国、柬埔寨、老挝、缅甸、锡金、尼泊尔、印度、斯里兰卡、马来西亚、印度尼西亚、菲律宾、日本（琉球）、澳大利亚和马达加斯加。模式标本采自印度尼西亚的爪哇岛。
本种为唇边组的一个常见种，其叶片长短与宽窄、叶柄的长短、乃至鳞片的颜色等均有一定的变异幅度，这些特征受环境的影响显著。分布到中国及邻近地区、曾被订名为 V. zosterifolia (Borry) Willd. 和 V. merrillii Christ 的标本均应为 V. elongata Sw. 本种同姬书带蕨的显著不同点在于形体大小上的差异，此外，二者的鳞片和孢子囊群中的隔丝也有明显的区别。
《Flora of China》 Vol. 2-3 (2013)
Haplopteris elongata (Swartz) E. H. Crane Syst. Bot. 22: 514. 1998.
唇边书带蕨 chun bian shu dai jue
Vittaria elongata Swartz, Syn. Fil. 109, 302. 1806; V. formosana Nakai; V. ogasawarensis Kodama; V. pauciariolata Ching; V. tortifrons Hayata.
Rhizome rather long creeping, much branched, bearing roots with very numerous water-absorbing root hairs; scales dark brown, brightly iridescent, subulate-lanceolate, 4-5 mm, 0.5-1 mm wide at base, margin prominently denticulate, apex bristlelike, areole wall thick, dark colored. Fronds clustered, drooping; stipe ca. 0.5 mm distant; lamina thinly leathery, linear or ribbonlike, up to 100 × 0.5-2 cm, base gradually narrowed, apex rounded or obtuse; costa slender, not prominent, veins evident. Soral line marginal, immersed in deep groove, open outward, fertile throughout; paraphyses many, long, with head obconic, longer than wide. Spores monolete, narrowly oblong in outline, surface ornamentation obscure.
Epiphytic or epilithic; 100-1400 m. Fujian, Guangdong, Guangxi, Hainan, Taiwan, SE Xizang, S Yunnan [Indonesia, S Japan, Laos, Malaysia, Myanmar, Nepal, Philippines, Sri Lanka, Thailand, Vietnam; Australia, Madagascar].
Haplopteris elongata always grows with other epiphytic ferns, notably with Pseudodrynaria coronans (Aglaomorpha coronans) and species of the Asplenium nidus and A. laserpitiifolium groups. It is a polymorphic species, with fluctuations in plant size and scale color depending on the habitat. All Chinese records of Vittaria zosterifolia Willdenow and V. merrillii Christ were based on misidentifications of this species.
别名：长叶书带蕨;
科名：书带蕨科 Vittariaceae
属名：书带蕨属 Vittaria
12. 唇边书带蕨（中国高等植物图鉴）长叶书带蕨（中国主要植物图说蕨类植物门）、阔叶书带蕨（中国蕨类植物图谱）、垂叶书带蕨（台湾植物志）、双唇书带蕨（西藏植物志）图版7：1-4.
Vittaria elongata Sw., Syn. Fil. 109, 302. 1806; Hook. et Bak. Syn. Fil. 395. 1874, pro parte; Bedd., Ferns South India t. 21. 1863 et Handb. Ferns Brit. India 404, f. 238. 1883; Clarke in Trans. Linn. Soc. Lond. II. Bot. 1: 573. 1880; Ching in Sinensia 1 (12): 179. 1931; H. Ito in Journ. Jap. Bot. 12: 466. 1936; Tard. -Blot et C. Chr. in Lecomte, Fl. Indo-Chine 7 (2): 197. 1940; Holtt., Rev. Fl. Mat. 2: 614, f. 360. 1955 et in Dansk Bot. Ark. 20: 34. 1961; 傅书遐, 中国主要植物图说 蕨类植物门 252, 图338. 1957; Bir in Res. Bull. (N. S.) Panjab Univ. 13 (1-2): 17, f. 1-5. 1962; Ching et al. in W. Y. Chun et al., Fl. Hainan. 1: 201. 1964; Ic. Corm. Sin. 1: 281, f. 561. 1972; DeVol in H. L. Li et al., Fl. Taiwan 1: 238. 1975; Y. L. Chang et al., Sporae Pterid. Sin. 375, t. 87, f. 5-6, 1976; Dixit in Journ. Econ. Tax. Bot. 2: 210, f. 1-9. 1981; Fl. Fujian. 1: 260, f. 246. 1982; Tagawa et K. Iwats., Fl. Thail. 3 (2): 223. 1985. ——Pteris graminifolia Roxb. in Calc. Journ. 4: 502, t. 33. 1844. ——Vittaria anodontolepis Fee, Mem. Foug. 3: 23, t. 4, f. 3. 1851-52. ——Vittaria boninensis Christ in Warb. Monsunia 1: 57. 1900. ——Vittaria ogasawarensis Kodama in Matsum., Ic. Pl. Koishikav. 3: 146. 1915; H. Ito, l. c. 467. 1936. ——Vittaria tortifrons Hayata, Ic. Pl. Forms. 6: 162. 1916; H. Ito, l. c. 466. 1936. ——Vittaria formosana Nakai in Bot. Mag. Tokyo 39: 176. 1925. ——Vittaria pauciareolata Ching in Sinensia 1 (1): 11. 1929 et in Hu et Ching, Ic. Fil. Sin. 1: pl. 38. 1930. ——Vittaria zosterifolia auct. non Willd. : Ching et al. in Chun et al., Fl. Hainan. 1: 202. 1964; Ic. Corm. Sin. 1: 281. 1972; DeVol in H. L. Li et al., Fl. Taiwan 1: 238. 1975; Y. L. Chang et al., Sporae Pterid. Sin. 377, t. 87, f. 7-8. 1976; Nakaike, Enum. Pterid. Jap. Fil. 84. 1975 et New Fl. Jap. Pterid. 716, cum f. 1992; Dixit, 1. c. 212, f. 8. 1981; Fl. Fujian. 1: 260. 1982; Ching et S. K. Wu in C. Y. Wu, Fl. Xizang. 1: 105, f. 26: 1-2, excl. 3. 1983; X. Cheng in W. T. Wang, Vasc. Pl. Hengduan Mount. 1: 64. 1993. ——Vittaria merrillii auct. non Christ: Ching, l. c. 180. 1931; Dixit, l. c. f. 9. 1981.
根状茎长而横走，多分叉，密被须根和鳞片；须根的根毛绒毛状，密布，形成线状的吸水结构；鳞片黑褐色，具亮光泽，钻状披针形，长4-5毫米，基部最宽，约0.5-1毫米，先端渐尖，纤毛状，端部呈腺体状，粗筛孔网眼壁增厚，黑色，边缘具较长的睫毛状齿；叶稍远生，相距约0.5毫米，通常成丛倒垂。叶柄横切面椭圆形，或长或短，基部常被较根状茎鳞片狭长的鳞片；叶片线形或带状，长可达1米以上，宽0.5-2厘米，顶端圆头或钝头，下部渐狭，全缘；中肋细，两面扁平，不甚明显，侧脉多数，形成斜升的网眼，较明显。叶近革质，干后有皱纹。孢子囊群线着生于叶缘的双唇状夹缝中，开口向外，自叶片的近基部延伸直达近顶端；隔丝多数，顶端杯状，长略大于宽。孢子狭长椭圆型，单裂缝，表面纹饰模糊。
分布于福建（永泰、宁德）、台湾（全岛分布）、海南（全岛分布）、广东（信宜、肇庆、阳春、阳江）、广西（南宁、龙州）、云南（河口、屏边、金平、勐腊、勐海、景洪、福贡、贡山）、西藏（墨脱）。附生于树上或林中岩石上，常同王冠蕨 Pseudodrynaria coronans (Wall. ex Mett.) Ching，鸟巢蕨属 Neottopteris 及铁角蕨属 Asplenium 等附生蕨类植物混生，海拔190-1350米。也分布于越南、泰国、柬埔寨、老挝、缅甸、锡金、尼泊尔、印度、斯里兰卡、马来西亚、印度尼西亚、菲律宾、日本（琉球）、澳大利亚和马达加斯加。模式标本采自印度尼西亚的爪哇岛。
本种为唇边组的一个常见种，其叶片长短与宽窄、叶柄的长短、乃至鳞片的颜色等均有一定的变异幅度，这些特征受环境的影响显著。分布到中国及邻近地区、曾被订名为 V. zosterifolia (Borry) Willd. 和 V. merrillii Christ 的标本均应为 V. elongata Sw. 本种同姬书带蕨的显著不同点在于形体大小上的差异，此外，二者的鳞片和孢子囊群中的隔丝也有明显的区别。
《Flora of China》 Vol. 2-3 (2013)
Haplopteris elongata (Swartz) E. H. Crane Syst. Bot. 22: 514. 1998.
唇边书带蕨 chun bian shu dai jue
Vittaria elongata Swartz, Syn. Fil. 109, 302. 1806; V. formosana Nakai; V. ogasawarensis Kodama; V. pauciariolata Ching; V. tortifrons Hayata.
Rhizome rather long creeping, much branched, bearing roots with very numerous water-absorbing root hairs; scales dark brown, brightly iridescent, subulate-lanceolate, 4-5 mm, 0.5-1 mm wide at base, margin prominently denticulate, apex bristlelike, areole wall thick, dark colored. Fronds clustered, drooping; stipe ca. 0.5 mm distant; lamina thinly leathery, linear or ribbonlike, up to 100 × 0.5-2 cm, base gradually narrowed, apex rounded or obtuse; costa slender, not prominent, veins evident. Soral line marginal, immersed in deep groove, open outward, fertile throughout; paraphyses many, long, with head obconic, longer than wide. Spores monolete, narrowly oblong in outline, surface ornamentation obscure.
Epiphytic or epilithic; 100-1400 m. Fujian, Guangdong, Guangxi, Hainan, Taiwan, SE Xizang, S Yunnan [Indonesia, S Japan, Laos, Malaysia, Myanmar, Nepal, Philippines, Sri Lanka, Thailand, Vietnam; Australia, Madagascar].
Haplopteris elongata always grows with other epiphytic ferns, notably with Pseudodrynaria coronans (Aglaomorpha coronans) and species of the Asplenium nidus and A. laserpitiifolium groups. It is a polymorphic species, with fluctuations in plant size and scale color depending on the habitat. All Chinese records of Vittaria zosterifolia Willdenow and V. merrillii Christ were based on misidentifications of this species.