密刺悬钩子Rubus subtibetanus Hand.-Mazz.
密刺悬钩子Rubus subtibetanus Hand.-Mazz.
35.密刺悬钩子 图版7: 1-3
Rubus subtibetanus Hand.-Mazz. in Sitzber. Math.-Nat. Cl. Akad. Wiss. Wien. 57: 268. 1920 et Symb. Sin. 7: 502. 1933. R. subtibetanus Hand.-Mazz. var. subtibetanus
形态特征：攀援灌木，扁1—2米；老枝褐色或红褐色，密被长短不等的针刺和短皮刺，并有柔毛、，小枝暗绿色，被较密针刺。小叶3—5枚，顶生小叶宽卵形至卵状披针形，顶端渐尖，基部截．形至微心形，比侧生小叶稍长，：边缘常羽状分裂，侧生小叶斜椭圆形或斜卵形，长2—5厘米，宽1．5—4厘米，顶端急尖至渐尖，基部圆形或宽楔形，上面有柔毛，下面密被灰白色至黄灰色绒毛，边缘有不整齐或缺刻状粗锯齿；叶柄长2．5一4.5厘米，顶生小叶柄长1．5—2．5厘米，侧生小叶近无柄，均被柔毛和较密针刺；托叶线状披针形，·具柔毛，长达1厘米。伞房花序顶生或腋生；总花梗和花梗均被柔毛和较密针刺；花梗长0．5一1厘米，至果期增长；苞片线形，具柔毛；花直径．6—8毫米；花萼外密被柔毛；萼片长卵形至卵状披针形，顶端急尖至渐尖，花时直立开展；果时常反折；花瓣近圆形，白色带红或紫红色；基部微具柔毛，稍长于萼片；雄蕊花丝线形，无毛；花柱稍长于雄蕊，无毛；子房具柔毛。果实近球形，直径6-8毫米，成熟时蓝黑色，微具柔毛；核较平滑或有细皱纹. 花期5—6月，果期6一7月。
产地分布：产陕西甜肃、四川.生山坡或山谷灌丛中,海拔2300米。
本种提示：此种和喜阴悬钩子R．Mesogaeus Focke：区别在于后者老枝具稀疏基部宽大的扁平皮；刺，小枝疏生针刺或近无刺；：顶生小叶宽菱状卵形或椭圆；果实无毛.又和西藏悬钩子-R.thibetanus Franch；区别在于后者枝上少刺，小叶较多（5）7-11枚，叶柄较短，顶生：小叶比侧生者长1倍以上，且常羽状深裂，均可区分。
亚组概述：Subsect．Stimuiantes：Yu et．Lu，．植物分类学报20(3)：299．1982．-Sect．Pungentes Focke，Bibl．Bot．72(2)：160；1911 et，83：40． 1914·P．P． -Sect。Ldaeanthi Focke，L c．72(2)：171.1911，et 83：·43．1914．P．P。 本亚组有下列13种。
《Flora of China》 Vol. 9 (2003)
Rubus subtibetanus Handel-Mazzetti
密刺悬钩子 mi ci xuan gou zi
Shrubs climbing, 1–2 m tall. Old branches brown or reddish brown, pubescent, with dense unequal needle-like prickles and basally broadened short prickles; branchlets dark green, more with dense, needle-like prickles. Leaves imparipinnate, 3–5-foliolate; petiole 2.5–4.5 cm, petiolule of terminal leaflet 1.5–2.5 cm, lateral leaflets subsessile, petiolules and rachis pubescent, with dense, needle-like prickles; stipules linear-lanceolate, 7–10 mm, pubescent; terminal leaflet broadly ovate to ovate-lanceolate, base truncate to subcordate, margin often pinnately lobed, apex acuminate, slightly longer than lateral leaflets, lateral leaflets oblique-elliptic or oblique-ovate, 2–5 × 1.5–4 cm, abaxially densely gray to yellowish gray tomentose, adaxially pubescent, margin irregularly or incised rough serrate, base rounded or broadly cuneate, apex acute to shortly acuminate. Inflorescences terminal or axillary, corymbose, 2–4 cm; rachis and pedicels pubescent, with dense, needle-like prickles; bracts linear, 5–7 mm, pubescent. Pedicel 6–10 mm. Flowers 6–8 mm in diam. Calyx abaxially densely pubescent; sepals erect at anthesis, reflexed in fruit, narrowly ovate to ovate-lanceolate, 5–7 × 2–3 mm, apex acute to acuminate. Petals white, with red or purplish red spots, suborbicular or obovate, 5–6 mm in diam., slightly longer than sepals, base puberulous. Stamens many, somewhat shorter than petals; filaments linear, glabrous. Pistils slightly longer than stamens; ovary pubescent; styles glabrous. Aggregate fruit blue-black at maturity, subglobose, 6–8 mm in diam., puberulous; pyrenes smooth or rugulose. Fl. May–Jun, fr. Jun–Jul.
Slopes, thickets, valleys; below 2300 m. Gansu, Shaanxi, Sichuan.
别名：康藏悬钩子;
科名：蔷薇科 Rosaceae
属名：悬钩子属 Rubus
35.密刺悬钩子 图版7: 1-3
Rubus subtibetanus Hand.-Mazz. in Sitzber. Math.-Nat. Cl. Akad. Wiss. Wien. 57: 268. 1920 et Symb. Sin. 7: 502. 1933. R. subtibetanus Hand.-Mazz. var. subtibetanus
形态特征：攀援灌木，扁1—2米；老枝褐色或红褐色，密被长短不等的针刺和短皮刺，并有柔毛、，小枝暗绿色，被较密针刺。小叶3—5枚，顶生小叶宽卵形至卵状披针形，顶端渐尖，基部截．形至微心形，比侧生小叶稍长，：边缘常羽状分裂，侧生小叶斜椭圆形或斜卵形，长2—5厘米，宽1．5—4厘米，顶端急尖至渐尖，基部圆形或宽楔形，上面有柔毛，下面密被灰白色至黄灰色绒毛，边缘有不整齐或缺刻状粗锯齿；叶柄长2．5一4.5厘米，顶生小叶柄长1．5—2．5厘米，侧生小叶近无柄，均被柔毛和较密针刺；托叶线状披针形，·具柔毛，长达1厘米。伞房花序顶生或腋生；总花梗和花梗均被柔毛和较密针刺；花梗长0．5一1厘米，至果期增长；苞片线形，具柔毛；花直径．6—8毫米；花萼外密被柔毛；萼片长卵形至卵状披针形，顶端急尖至渐尖，花时直立开展；果时常反折；花瓣近圆形，白色带红或紫红色；基部微具柔毛，稍长于萼片；雄蕊花丝线形，无毛；花柱稍长于雄蕊，无毛；子房具柔毛。果实近球形，直径6-8毫米，成熟时蓝黑色，微具柔毛；核较平滑或有细皱纹. 花期5—6月，果期6一7月。
产地分布：产陕西甜肃、四川.生山坡或山谷灌丛中,海拔2300米。
本种提示：此种和喜阴悬钩子R．Mesogaeus Focke：区别在于后者老枝具稀疏基部宽大的扁平皮；刺，小枝疏生针刺或近无刺；：顶生小叶宽菱状卵形或椭圆；果实无毛.又和西藏悬钩子-R.thibetanus Franch；区别在于后者枝上少刺，小叶较多（5）7-11枚，叶柄较短，顶生：小叶比侧生者长1倍以上，且常羽状深裂，均可区分。
亚组概述：Subsect．Stimuiantes：Yu et．Lu，．植物分类学报20(3)：299．1982．-Sect．Pungentes Focke，Bibl．Bot．72(2)：160；1911 et，83：40． 1914·P．P． -Sect。Ldaeanthi Focke，L c．72(2)：171.1911，et 83：·43．1914．P．P。 本亚组有下列13种。
《Flora of China》 Vol. 9 (2003)
Rubus subtibetanus Handel-Mazzetti
密刺悬钩子 mi ci xuan gou zi
Shrubs climbing, 1–2 m tall. Old branches brown or reddish brown, pubescent, with dense unequal needle-like prickles and basally broadened short prickles; branchlets dark green, more with dense, needle-like prickles. Leaves imparipinnate, 3–5-foliolate; petiole 2.5–4.5 cm, petiolule of terminal leaflet 1.5–2.5 cm, lateral leaflets subsessile, petiolules and rachis pubescent, with dense, needle-like prickles; stipules linear-lanceolate, 7–10 mm, pubescent; terminal leaflet broadly ovate to ovate-lanceolate, base truncate to subcordate, margin often pinnately lobed, apex acuminate, slightly longer than lateral leaflets, lateral leaflets oblique-elliptic or oblique-ovate, 2–5 × 1.5–4 cm, abaxially densely gray to yellowish gray tomentose, adaxially pubescent, margin irregularly or incised rough serrate, base rounded or broadly cuneate, apex acute to shortly acuminate. Inflorescences terminal or axillary, corymbose, 2–4 cm; rachis and pedicels pubescent, with dense, needle-like prickles; bracts linear, 5–7 mm, pubescent. Pedicel 6–10 mm. Flowers 6–8 mm in diam. Calyx abaxially densely pubescent; sepals erect at anthesis, reflexed in fruit, narrowly ovate to ovate-lanceolate, 5–7 × 2–3 mm, apex acute to acuminate. Petals white, with red or purplish red spots, suborbicular or obovate, 5–6 mm in diam., slightly longer than sepals, base puberulous. Stamens many, somewhat shorter than petals; filaments linear, glabrous. Pistils slightly longer than stamens; ovary pubescent; styles glabrous. Aggregate fruit blue-black at maturity, subglobose, 6–8 mm in diam., puberulous; pyrenes smooth or rugulose. Fl. May–Jun, fr. Jun–Jul.
Slopes, thickets, valleys; below 2300 m. Gansu, Shaanxi, Sichuan.