湖南悬钩子Rubus hunanensis Hand.-Mazz.
湖南悬钩子Rubus hunanensis Hand.-Mazz.
135.湖南悬钩子（拉汉种子植物名称）图版22：4-7
Rubus hunanensis Hand. -Mazz. Symb. Sin. 7: 497. f. 16. 1933: Migo in Journ. Shangh. Sci. Inst. Sect. 3 (4): 150. 1939. -R. buergeri Miq. var. viridifolius Hand.-Mazz. Symb. Sin. 7: 497. 1933. syn. nov.
攀援小灌木，高0.3-2米；枝细，密被细短柔毛，疏生钩状小皮刺。单叶，近圆形或宽卵形，直径8-13厘米，顶端急尖，基部深心形，幼时上面具细短柔毛，下面有绒毛和细柔毛，但绒毛逐渐脱落，老时两面近无毛，边缘5-7浅裂，裂片顶端急尖，稀圆钝，有不整齐锐锯齿，基部具掌状5出脉，侧脉2-3对；叶柄长6-9厘米，密被细短柔毛和稀疏钩状小皮刺；托叶离生，褐色，长达1厘米，不育枝上托叶长达1.8厘米，近掌状或羽状分裂，裂片线形，具细短柔毛，脱落或部分宿存。花数朵生于叶腋或成顶生短总状花序；总花梗和花梗密被灰色细短柔毛；花梗长0.5-1厘米；苞片与托叶相似；花直径0.7-1厘米；花萼外密被灰白色至黄灰色短柔毛和绒毛；萼片宽卵形，顶端急尖至短渐尖，外萼片较宽大，边缘羽状条裂，裂片线状披针形，内萼片较小，常不分裂，花后直立；花瓣倒卵形，白色，无毛；雄蕊短，花丝线形，无毛；雌蕊几与雄蕊等长，无毛。果实半球形，黄红色，包藏在宿萼内，无毛；核具细皱纹。花期7-8月，果期9-10月。
产江西、浙江、湖南、湖北、福建、台湾、广东、广西、四川、贵州。生山谷、山沟、密林或草丛中，海拔500-2500米。
此种和寒莓R. buergeri Miq.近似，但后者全体被绒毛状长柔毛；叶片的裂片顶端圆钝；花萼外密被淡黄色长柔毛和绒毛；萼片披针形或卵状披针形，外萼片较狭，仅顶端浅裂，易于区别。
形态特征：攀援小灌木，高0．3—2米；枝细，密被细短柔毛，疏生钩状小皮刺。单叶，近圆形或宽 卵形，直径8—13厘米，顶端急尖，基部深心形，幼时上面具细短柔毛，下面有绒毛和细柔 毛，但绒毛逐渐脱落，老时两面近无毛，边缘5一7浅裂；裂片顶端急尖，稀圆钝，有不整齐 锐锯齿，基部具掌状5出脉，侧脉2—3对；叶柄长6-9厘米，密被细短柔毛和稀疏钩状小 皮刺；托叶离生，褐色，长达1厘米，不育枝上托叶长达1．8厘米，近掌状或羽状分裂，裂片 线形，具细短柔毛，脱落或部分宿存。花数朵生于叶腋或成顶生短总状花序；总花梗和花 梗密被灰色细短柔毛；花梗长0．5—1厘米；苞片与托叶相似；花直径0．7—1厘米；花萼外 密被灰白色至黄灰色短柔毛和绒毛；萼片宽卵形，顶端急尖至短渐尖，外萼片较宽大，边缘 羽状条裂，裂片线状披针形，内萼片较小，常不分裂，花后直立；花瓣倒卵形，白色，无毛；雄 蕊短，花丝线形，无毛；雌蕊几与雄蕊等长，无毛。果实半球形，黄红色，包藏在宿萼内，无 毛；核具细皱纹。 花期?一8月，果期9—10月.
产地分布：产江西、浙江、湖南、湖北、福建、台湾、广东、广西、四川、贵州。生山谷、山沟、 密林或茸丛中，海拔500—2500米。
本种提示：此种和寒莓R．Buergeri Miq．近似，但后者全体被绒毛状长柔毛；叶片的裂片顶端圆钝；花萼外密被淡黄色长柔毛和绒毛；萼片披针形或卵状披针形，外萼片较狭，仅顶端浅 裂，易于区别。
参考文献：Rubus hunanensis Hand．—Mazz，Symb．Sin．7：497．F．16．1933；Migo in Journ．Shangh．Scl lnst．Sect．3(4)：150．1939．———R．Buergeri Miq·var．Viridi／olius Hand． Mazz．Symb．Sin．7：497．1933．Syn．Nov ．
亚组概述：Subsect．Moluccani(Focke) Yu et Lu,植物分类学报20(4)： 456．1982；，- Sect．Moluccami Focke，BiN．Pot．72(1)：71．1910 et 83：28．1914.P．P．
《Flora of China》 Vol. 9 (2003)
Rubus hunanensis Handel-Mazzetti
湖南悬钩子 hu nan xuan gou zi
Rubus buergeri Miquel var. viridifolius Handel-Mazzetti.
Shrubs climbing, 0.3–2 m tall. Branchlets slender, densely thinly pubescent, with sparse, curved minute prickles. Leaves simple; petiole 6–9 cm, densely thinly pubescent, with sparse, curved minute prickles; stipules free, brown, to 1 cm, stipules caducous or partly persistent, on sterile branchlets to 1.8 cm, nearly palmately or pinnately lobed, lobes linear, thinly pubescent; blade suborbicular or broadly ovate, 8–13 cm in diam., palmately 5-veined, lateral veins 2 or 3 pairs, raised abaxially, abaxially tomentose, intermixed thinly pubescent, hairs gradually deciduous, adaxially thinly pubescent when young, both surfaces subglabrescent, base deeply cordate, margin 5–7-lobed; lobes unevenly sharply serrate, apex acute, rarely obtuse. Inflorescences terminal, short subracemes, or several flowers in leaf axils; rachis and pedicels densely gray thinly pubescent; bracts similar to stipules, but smaller. Pedicel 5–10 mm. Flowers 7–10 mm in diam. Calyx abaxially densely gray to yellowish gray pubescent and tomentose; sepals erect after anthesis, broadly ovate, 6–8 × 4–7 mm, apex acute to shortly acuminate; outer sepals larger, pinnately laciniate, lobes linear-lanceolate, inner sepals smaller, usually undivided. Petals white, obovate, glabrous. Stamens shorter; filaments linear, glabrous. Pistils nearly as long as stamens, glabrous. Aggregate fruit yellowish red, semiglobose, glabrous, enclosed in persistent calyx; pyrenes rugulose. Fl. Jul–Aug, fr. Sep–Oct.
Montane valleys, forests, grasslands; 300--2500 m. Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Taiwan, Zhejiang.
Further study is required to determine the identity of Rubus pseudoacer Makino var. flexuosus Y. C. Liu & F. Y. Lu (Quart. J. Chin. Forest. 9(2): 133. 1976; R. pseudoacer subsp. flexuosus (T. S. Liu & F. Y. Lu) H. Ohashi & C. F. Hsieh), described from Taiwan, which might be synonymous with R. hunanensis.
别名：湖南莓;
科名：蔷薇科 Rosaceae
属名：悬钩子属 Rubus
135.湖南悬钩子（拉汉种子植物名称）图版22：4-7
Rubus hunanensis Hand. -Mazz. Symb. Sin. 7: 497. f. 16. 1933: Migo in Journ. Shangh. Sci. Inst. Sect. 3 (4): 150. 1939. -R. buergeri Miq. var. viridifolius Hand.-Mazz. Symb. Sin. 7: 497. 1933. syn. nov.
攀援小灌木，高0.3-2米；枝细，密被细短柔毛，疏生钩状小皮刺。单叶，近圆形或宽卵形，直径8-13厘米，顶端急尖，基部深心形，幼时上面具细短柔毛，下面有绒毛和细柔毛，但绒毛逐渐脱落，老时两面近无毛，边缘5-7浅裂，裂片顶端急尖，稀圆钝，有不整齐锐锯齿，基部具掌状5出脉，侧脉2-3对；叶柄长6-9厘米，密被细短柔毛和稀疏钩状小皮刺；托叶离生，褐色，长达1厘米，不育枝上托叶长达1.8厘米，近掌状或羽状分裂，裂片线形，具细短柔毛，脱落或部分宿存。花数朵生于叶腋或成顶生短总状花序；总花梗和花梗密被灰色细短柔毛；花梗长0.5-1厘米；苞片与托叶相似；花直径0.7-1厘米；花萼外密被灰白色至黄灰色短柔毛和绒毛；萼片宽卵形，顶端急尖至短渐尖，外萼片较宽大，边缘羽状条裂，裂片线状披针形，内萼片较小，常不分裂，花后直立；花瓣倒卵形，白色，无毛；雄蕊短，花丝线形，无毛；雌蕊几与雄蕊等长，无毛。果实半球形，黄红色，包藏在宿萼内，无毛；核具细皱纹。花期7-8月，果期9-10月。
产江西、浙江、湖南、湖北、福建、台湾、广东、广西、四川、贵州。生山谷、山沟、密林或草丛中，海拔500-2500米。
此种和寒莓R. buergeri Miq.近似，但后者全体被绒毛状长柔毛；叶片的裂片顶端圆钝；花萼外密被淡黄色长柔毛和绒毛；萼片披针形或卵状披针形，外萼片较狭，仅顶端浅裂，易于区别。
形态特征：攀援小灌木，高0．3—2米；枝细，密被细短柔毛，疏生钩状小皮刺。单叶，近圆形或宽 卵形，直径8—13厘米，顶端急尖，基部深心形，幼时上面具细短柔毛，下面有绒毛和细柔 毛，但绒毛逐渐脱落，老时两面近无毛，边缘5一7浅裂；裂片顶端急尖，稀圆钝，有不整齐 锐锯齿，基部具掌状5出脉，侧脉2—3对；叶柄长6-9厘米，密被细短柔毛和稀疏钩状小 皮刺；托叶离生，褐色，长达1厘米，不育枝上托叶长达1．8厘米，近掌状或羽状分裂，裂片 线形，具细短柔毛，脱落或部分宿存。花数朵生于叶腋或成顶生短总状花序；总花梗和花 梗密被灰色细短柔毛；花梗长0．5—1厘米；苞片与托叶相似；花直径0．7—1厘米；花萼外 密被灰白色至黄灰色短柔毛和绒毛；萼片宽卵形，顶端急尖至短渐尖，外萼片较宽大，边缘 羽状条裂，裂片线状披针形，内萼片较小，常不分裂，花后直立；花瓣倒卵形，白色，无毛；雄 蕊短，花丝线形，无毛；雌蕊几与雄蕊等长，无毛。果实半球形，黄红色，包藏在宿萼内，无 毛；核具细皱纹。 花期?一8月，果期9—10月.
产地分布：产江西、浙江、湖南、湖北、福建、台湾、广东、广西、四川、贵州。生山谷、山沟、 密林或茸丛中，海拔500—2500米。
本种提示：此种和寒莓R．Buergeri Miq．近似，但后者全体被绒毛状长柔毛；叶片的裂片顶端圆钝；花萼外密被淡黄色长柔毛和绒毛；萼片披针形或卵状披针形，外萼片较狭，仅顶端浅 裂，易于区别。
参考文献：Rubus hunanensis Hand．—Mazz，Symb．Sin．7：497．F．16．1933；Migo in Journ．Shangh．Scl lnst．Sect．3(4)：150．1939．———R．Buergeri Miq·var．Viridi／olius Hand． Mazz．Symb．Sin．7：497．1933．Syn．Nov ．
亚组概述：Subsect．Moluccani(Focke) Yu et Lu,植物分类学报20(4)： 456．1982；，- Sect．Moluccami Focke，BiN．Pot．72(1)：71．1910 et 83：28．1914.P．P．
《Flora of China》 Vol. 9 (2003)
Rubus hunanensis Handel-Mazzetti
湖南悬钩子 hu nan xuan gou zi
Rubus buergeri Miquel var. viridifolius Handel-Mazzetti.
Shrubs climbing, 0.3–2 m tall. Branchlets slender, densely thinly pubescent, with sparse, curved minute prickles. Leaves simple; petiole 6–9 cm, densely thinly pubescent, with sparse, curved minute prickles; stipules free, brown, to 1 cm, stipules caducous or partly persistent, on sterile branchlets to 1.8 cm, nearly palmately or pinnately lobed, lobes linear, thinly pubescent; blade suborbicular or broadly ovate, 8–13 cm in diam., palmately 5-veined, lateral veins 2 or 3 pairs, raised abaxially, abaxially tomentose, intermixed thinly pubescent, hairs gradually deciduous, adaxially thinly pubescent when young, both surfaces subglabrescent, base deeply cordate, margin 5–7-lobed; lobes unevenly sharply serrate, apex acute, rarely obtuse. Inflorescences terminal, short subracemes, or several flowers in leaf axils; rachis and pedicels densely gray thinly pubescent; bracts similar to stipules, but smaller. Pedicel 5–10 mm. Flowers 7–10 mm in diam. Calyx abaxially densely gray to yellowish gray pubescent and tomentose; sepals erect after anthesis, broadly ovate, 6–8 × 4–7 mm, apex acute to shortly acuminate; outer sepals larger, pinnately laciniate, lobes linear-lanceolate, inner sepals smaller, usually undivided. Petals white, obovate, glabrous. Stamens shorter; filaments linear, glabrous. Pistils nearly as long as stamens, glabrous. Aggregate fruit yellowish red, semiglobose, glabrous, enclosed in persistent calyx; pyrenes rugulose. Fl. Jul–Aug, fr. Sep–Oct.
Montane valleys, forests, grasslands; 300--2500 m. Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Taiwan, Zhejiang.
Further study is required to determine the identity of Rubus pseudoacer Makino var. flexuosus Y. C. Liu & F. Y. Lu (Quart. J. Chin. Forest. 9(2): 133. 1976; R. pseudoacer subsp. flexuosus (T. S. Liu & F. Y. Lu) H. Ohashi & C. F. Hsieh), described from Taiwan, which might be synonymous with R. hunanensis.