华南悬钩子Rubus hanceanus Ktze.
华南悬钩子Rubus hanceanus Ktze.
156.华南悬钩子 图版25:3-4
Rubus hanceanus Ktze. Meth. Sp. Rub. 72 et 77. 1879; Forbes et Hemsl. in Journ. Linn. Soc. Bot. 23: 231. 1887. ——R. fordii Hance in Journ. Bot. 21: 298. 1883. ——R. hirtiflorus Card. in Lecomte, Not. Syst. 3: 290. 1914. ——R. prandianus Hand. -Mazz. in Sitzber. Math. -Nat. Cl. Acad. Wiss. Wien. 58: 91. 1921 et Symb. Sin. 7: 487. 1933.
藤状或攀援小灌木，高约1米；枝密被灰白色绒毛，老时逐渐脱落，具稀疏钩状小皮刺或有腺毛。单叶，非革质，心状宽卵形，长6-11厘米，宽4-8厘米，顶端渐尖，基部深心形，上面深褐色，仅于叶脉具柔毛，下面密被灰白色或浅黄灰色绒毛，叶脉5-7对，边缘浅裂，有不整齐锐锯齿；叶柄长1-2厘米，幼时具灰白色绒毛，逐渐脱落，有稀疏小皮刺；托叶早落。顶生总状花序，较长大，有少数花；总花梗、花梗和花萼均密被长2-4毫米的腺毛和绒毛状长柔毛，并疏生针刺；花梗长1.5-2.5厘米；苞片与托叶相似，长圆形或椭圆形，全缘或顶端浅条裂，有长柔毛和疏腺毛，膜质，早落；花直径1-1.5厘米；花萼外密被紫褐色较长腺毛和针刺；萼片宽卵形，长10-12毫米，顶端尾尖，全缘，在果期常.直立开展，稀反折；花瓣宽椭圆形，未开时红色，短于萼片，具柔毛，基部有短爪；雄蕊花丝宽扁而短，花药稍具长柔毛；花柱长于雄蕊很多，子房无毛。果实近球形，直径1-1.5厘米，黑色，无毛；小核果半圆形或近肾形；核稍具皱纹。花期3-5月，果期6-7月。
产湖南、广东、广西、福建。普遍生低海拔的山谷疏林、竹林下或岩石阴处。
形态特征：藤状或攀援小灌木，高约1米；枝密被灰白色绒毛，老时逐渐脱落，具稀疏钩状小皮刺或有腺毛。单叶，非革质，状宽卵形，长6—11厘米，宽斗一8厘米，顶端渐尖，基部深心形，上面深褐色，仅于叶脉具柔毛，下面密被灰白色或浅黄灰色绒毛，叶脉5一7对，边缘浅裂，有不整齐锐锯齿；叶柄长1—2厘米，幼时具灰白色绒毛，逐渐脱落，有稀疏小皮刺；托叶早落。顶生总状花序，较长大，有少数花；总花梗、花梗和花萼均密被长2—4毫米的腺毛和绒毛状长柔毛，并疏生针刺；花梗长1．5—2．5厘米；苞片与托叶相似，长圆形或椭圆形，全缘或顶端浅条裂，有长柔毛和疏腺毛，膜质，早落；花直径1—1．5厘米；花萼外密被紫褐色较长腺毛和针刺；片宽卵形，长19—12毫米，顶端尾尖，全缘，在果期常直立开展，稀反折；花瓣宽椭圆形，未开时红色，短于萼片，具柔毛，基部有短爪；雄蕊花丝宽扁而短，花药稍具长柔毛；花柱长于雄蕊很多，子房无毛。果实近球形；直径1-1.5厘米；黑色，无毛；小核果半圆形或近肾形；核稍具皱纹。 花期3—5月，果期6一7月。
产地分布：产湖南、广东、广西、福建。普遍生低海拔的山谷疏林、竹林下或岩石阴处。
参考文献：Rubus hanceanus Ktze·．Meth．SP．Rub：72 et 7．1879；Forbes et HemsI．In Journ．Linn，Soc.Bot．23：231．1887．——R. fordii Hance in Journ．Bot．21：298．1883．——R．Hirtiflorus card in in Lecomte，Not。Syst，3：290．1914． —R，prandianus Hand．—Mazz．InSitzber·，Math．—Nat．Cl．Acad．Wiss．Wien．58：91．192l ec Symb．Sin。?：487．1933．
亚组概述：Subsect；Sozostyli；(Focke)yu et lu，植物分类学报20(4)：461．1982．-Sect．Sozostyli Focke，Bibl．Bot。—72(1):42.1910 et 83:22. 1914. 本亚组有下列17种。
《Flora of China》 Vol. 9 (2003)
Rubus hanceanus Kuntze
华南悬钩子 hua nan xuan gou zi
Rubus fordii Hance; R. hirtiflorus Cardot; R. prandianus Handel-Mazzetti.
Shrubs lianoid or scandent, ca. 1 m tall. Branchlets brown or reddish brown, terete, densely gray tomentose, glabrescent, with sparse, curved minute prickles or stipitate glands. Leaves simple; petiole 1–2 cm, gray tomentose at first, gradually glabrescent, with sparse, minute prickles; stipules early caducous, free, oblong or elliptic, villous, stipitate glandular, margin entire or apically laciniate; blade broadly ovate, 6–11 × 4–8 cm; veins 5–7 pairs, prominent abaxially, somewhat impressed adaxially, not leathery, abaxially densely gray or yellowish gray tomentose, adaxially dark brown, glabrous except soft hairy along veins, base cordate, margin lobed, unevenly sharply serrate, apex acuminate. Inflorescences terminal, racemes, 6–9 cm, loosely few flowered; rachis and pedicels tomentose-villous, with stipitate glands 2–4 mm and sparse, needle-like prickles; bracts caducous, oblong or elliptic, 5–8 mm, membranous, villous, with sparse, intermixed stipitate glandular hairs, margin entire or apically lobed. Pedicel 1.5–2.5 cm. Flowers 1–1.5 cm in diam. Calyx abaxially densely tomentose-villous, with purplish brown long stalked glands and needle-like prickles; sepals erect, rarely reflexed in fruit, broadly ovate, 1–1.2 cm × 4–6 mm, margin entire, apex caudate. Petals broadly elliptic, 8–11 × 5–7 mm, soft hairy, base shortly clawed, apex obtuse. Stamens many, shorter than petals; filaments short, broad, complanate; anthers slightly villous. Pistils much longer than stamens; style and ovary glabrous. Aggregate fruit black, subglobose, 1–1.5 cm in diam., glabrous; drupelets subreniform or semiglobose; pyrenes somewhat rugose. Fl. Mar–May, fr. Jun–Jul.
Sparse mixed forests in valleys, bamboo thickets, shaded rocky places; Low elevations. Fujian, Guangdong, Guangxi, Hunan.
别名：
科名：蔷薇科 Rosaceae
属名：悬钩子属 Rubus
156.华南悬钩子 图版25:3-4
Rubus hanceanus Ktze. Meth. Sp. Rub. 72 et 77. 1879; Forbes et Hemsl. in Journ. Linn. Soc. Bot. 23: 231. 1887. ——R. fordii Hance in Journ. Bot. 21: 298. 1883. ——R. hirtiflorus Card. in Lecomte, Not. Syst. 3: 290. 1914. ——R. prandianus Hand. -Mazz. in Sitzber. Math. -Nat. Cl. Acad. Wiss. Wien. 58: 91. 1921 et Symb. Sin. 7: 487. 1933.
藤状或攀援小灌木，高约1米；枝密被灰白色绒毛，老时逐渐脱落，具稀疏钩状小皮刺或有腺毛。单叶，非革质，心状宽卵形，长6-11厘米，宽4-8厘米，顶端渐尖，基部深心形，上面深褐色，仅于叶脉具柔毛，下面密被灰白色或浅黄灰色绒毛，叶脉5-7对，边缘浅裂，有不整齐锐锯齿；叶柄长1-2厘米，幼时具灰白色绒毛，逐渐脱落，有稀疏小皮刺；托叶早落。顶生总状花序，较长大，有少数花；总花梗、花梗和花萼均密被长2-4毫米的腺毛和绒毛状长柔毛，并疏生针刺；花梗长1.5-2.5厘米；苞片与托叶相似，长圆形或椭圆形，全缘或顶端浅条裂，有长柔毛和疏腺毛，膜质，早落；花直径1-1.5厘米；花萼外密被紫褐色较长腺毛和针刺；萼片宽卵形，长10-12毫米，顶端尾尖，全缘，在果期常.直立开展，稀反折；花瓣宽椭圆形，未开时红色，短于萼片，具柔毛，基部有短爪；雄蕊花丝宽扁而短，花药稍具长柔毛；花柱长于雄蕊很多，子房无毛。果实近球形，直径1-1.5厘米，黑色，无毛；小核果半圆形或近肾形；核稍具皱纹。花期3-5月，果期6-7月。
产湖南、广东、广西、福建。普遍生低海拔的山谷疏林、竹林下或岩石阴处。
形态特征：藤状或攀援小灌木，高约1米；枝密被灰白色绒毛，老时逐渐脱落，具稀疏钩状小皮刺或有腺毛。单叶，非革质，状宽卵形，长6—11厘米，宽斗一8厘米，顶端渐尖，基部深心形，上面深褐色，仅于叶脉具柔毛，下面密被灰白色或浅黄灰色绒毛，叶脉5一7对，边缘浅裂，有不整齐锐锯齿；叶柄长1—2厘米，幼时具灰白色绒毛，逐渐脱落，有稀疏小皮刺；托叶早落。顶生总状花序，较长大，有少数花；总花梗、花梗和花萼均密被长2—4毫米的腺毛和绒毛状长柔毛，并疏生针刺；花梗长1．5—2．5厘米；苞片与托叶相似，长圆形或椭圆形，全缘或顶端浅条裂，有长柔毛和疏腺毛，膜质，早落；花直径1—1．5厘米；花萼外密被紫褐色较长腺毛和针刺；片宽卵形，长19—12毫米，顶端尾尖，全缘，在果期常直立开展，稀反折；花瓣宽椭圆形，未开时红色，短于萼片，具柔毛，基部有短爪；雄蕊花丝宽扁而短，花药稍具长柔毛；花柱长于雄蕊很多，子房无毛。果实近球形；直径1-1.5厘米；黑色，无毛；小核果半圆形或近肾形；核稍具皱纹。 花期3—5月，果期6一7月。
产地分布：产湖南、广东、广西、福建。普遍生低海拔的山谷疏林、竹林下或岩石阴处。
参考文献：Rubus hanceanus Ktze·．Meth．SP．Rub：72 et 7．1879；Forbes et HemsI．In Journ．Linn，Soc.Bot．23：231．1887．——R. fordii Hance in Journ．Bot．21：298．1883．——R．Hirtiflorus card in in Lecomte，Not。Syst，3：290．1914． —R，prandianus Hand．—Mazz．InSitzber·，Math．—Nat．Cl．Acad．Wiss．Wien．58：91．192l ec Symb．Sin。?：487．1933．
亚组概述：Subsect；Sozostyli；(Focke)yu et lu，植物分类学报20(4)：461．1982．-Sect．Sozostyli Focke，Bibl．Bot。—72(1):42.1910 et 83:22. 1914. 本亚组有下列17种。
《Flora of China》 Vol. 9 (2003)
Rubus hanceanus Kuntze
华南悬钩子 hua nan xuan gou zi
Rubus fordii Hance; R. hirtiflorus Cardot; R. prandianus Handel-Mazzetti.
Shrubs lianoid or scandent, ca. 1 m tall. Branchlets brown or reddish brown, terete, densely gray tomentose, glabrescent, with sparse, curved minute prickles or stipitate glands. Leaves simple; petiole 1–2 cm, gray tomentose at first, gradually glabrescent, with sparse, minute prickles; stipules early caducous, free, oblong or elliptic, villous, stipitate glandular, margin entire or apically laciniate; blade broadly ovate, 6–11 × 4–8 cm; veins 5–7 pairs, prominent abaxially, somewhat impressed adaxially, not leathery, abaxially densely gray or yellowish gray tomentose, adaxially dark brown, glabrous except soft hairy along veins, base cordate, margin lobed, unevenly sharply serrate, apex acuminate. Inflorescences terminal, racemes, 6–9 cm, loosely few flowered; rachis and pedicels tomentose-villous, with stipitate glands 2–4 mm and sparse, needle-like prickles; bracts caducous, oblong or elliptic, 5–8 mm, membranous, villous, with sparse, intermixed stipitate glandular hairs, margin entire or apically lobed. Pedicel 1.5–2.5 cm. Flowers 1–1.5 cm in diam. Calyx abaxially densely tomentose-villous, with purplish brown long stalked glands and needle-like prickles; sepals erect, rarely reflexed in fruit, broadly ovate, 1–1.2 cm × 4–6 mm, margin entire, apex caudate. Petals broadly elliptic, 8–11 × 5–7 mm, soft hairy, base shortly clawed, apex obtuse. Stamens many, shorter than petals; filaments short, broad, complanate; anthers slightly villous. Pistils much longer than stamens; style and ovary glabrous. Aggregate fruit black, subglobose, 1–1.5 cm in diam., glabrous; drupelets subreniform or semiglobose; pyrenes somewhat rugose. Fl. Mar–May, fr. Jun–Jul.
Sparse mixed forests in valleys, bamboo thickets, shaded rocky places; Low elevations. Fujian, Guangdong, Guangxi, Hunan.