阔叶瓦韦Lepisorus tosaensis (Makino) H. Ito
阔叶瓦韦Lepisorus tosaensis (Makino) H. Ito
8.阔叶瓦韦（中国高等植物图鉴）拟瓦韦（台湾植物志）图版10：1-2
Lepisorus tosaensis (Makino) H. Ito in J. Jap. Bot. 11: 93. 1935; Pic Ser., Ind. Fil. Sup. 4: 177. 1965; Tagawa, Col. Illustr. Jap. Pterid. 158. Pl. 67-363. 1959; Y. L. Zhang et al., Sporae Pterid. Sin. 493. 1976; De Vol et C. M. Kuo in H. L. Li et al., Fl. Taiwan 1: 188. 1975 et Fl. Taiwan 2nd ed. 1: 493. 1994; Nakaike, Illustr. Pterid. Jap. 2: 444. f. 445. 1981; Fl. Fujian 1: 230. 1982; 陈仁钧, 安徽植物志1: 189. 图195. 1985; S. X. Xu in j, F. Cheng et G. F. Chu, Fl. Jiangxi 1: 313. 1993; Z. F. Zhang et S. Y. Zhang, Fl. Zhejiang 1: 317. f. 334. 1992; S. L. Yu et al. in Bull. Bot. Res. 16 (1): 10. 1996. ——Polypodium tosaense Makino in Bot. Mag. Tokyo 27: 127. 1913. ——Polypodium lineare Thunb. var. caudatum Makino in Bot. Mag. Tokyo 17: 78. 1903. ——Polypodium infraplanicostale Hayata, Icon. Pl. Form. 5: 315. f. 128. 1915. ——Lepisorus infraplanicostalis (Hayata) Ching in Bull. Fan Mem. Inst. Biol. Bot. 4: 92. 1933. ——Polypodium morii Hayata, Pl. Form. 8: 153. f. 83, 84. 1919. ——Polypodium haozanense Hayata, Fl. Form. 8: 152. Pl. 81, 82. 1919.
植株高约15-30厘米。根状茎短促横卧，密被卵状披针形鳞片；鳞片深棕色，大部分不透明，仅边缘有1-2行淡棕色透明的细胞。叶簇生或近生；叶柄长1-5厘米，禾秆色；叶片披针形，中部为最宽约1-2厘米，向两端渐变狭，顶端渐尖头，基部渐狭并下延，长 (10) 13-20厘米，干后淡棕色或灰绿色，革质，两面光滑无毛。主脉上下均隆起，小脉不见。孢子囊群圆形，位于主脉与叶缘之间，聚生于叶片上半部，幼时被淡棕色圆形的隔丝覆盖。
产浙江 (临安、鄞县、开化、丽水、遂昌、庆云、平阳、宁波、杭州和苍南) 、江西 (遂川、宜丰、修水、寻乌) 、安徽 (黟县) , 福建 (建德、德化、南平、崇安) 、台湾 (南投) 、广东 (增城南昆山、翁源至英德、乐昌、状元山、乳源、新兴、信宜、河源、惠阳) 、香港、广西 (龙胜、武鸣、大明山、平南、瑶山) 、云南 (昆明、墨江、勐海、屏边、顺宁、金平、河口、文山) 、贵州 (江口、梵净山、安顺、凯里) 、四川 (峨眉山、会里、屏山) 、重庆 (北碚) 、湖南 (黔阳、莽山、保靖) 、西藏 (墨脱) 。附生溪边林下树干或岩石上, 及石灰墙缝中, 海拔650-1700米。模式标本采自日本。
《Flora of China》 Vol. 2-3 (2013)
Lepisorus tosaensis (Makino) H. It? J. Jap. Bot. 11: 93. 1935.
阔叶瓦韦 kuo ye wa wei
Polypodium tosaense Makino, Bot. Mag. (Tokyo) 27: 127. 1913; Lepisorus infraplanicostalus (Hayata) Ching; L. paohuashanensis Ching; L. sinicus Ching & Z. Y. Liu; Polypodium hoozanense Hayata; P. infraplanicostale Hayata; P. lineare N. L. Burman var. caudatum Makino; P. morii Hayata.
Plants 15-30 cm tall. Rhizomes short and decumbent, less than 3 cm, 2-3 mm in diam., apex densely scaly, remainder soon naked; scales basifixed, deep brown with paler margins, iridescent when young, lanceolate or broadly lanceolate, 2-3.5 × 0.7-1.1 mm; most lumina opaque, marginal 1 or 2 rows of cells brownish, transparent. Fronds clustered; stipe straw-colored, 1-3(-5) cm, 1-1.5 mm in diam.; lamina brownish or grayish green when dried, lanceolate or linear-lanceolate, both ends attenuate, (10-)13-20 × 1-2 cm, widest at middle, leathery, both surfaces glabrous, base cuneate, decurrent, apex acuminate; costa raised on both sides, veinlets obscure. Sori on distal half of lamina, costular, rarely midway between costa and margins, orbicular, 3-7 mm in diam.; paraphyses orbicular, brownish with central lumina small and opaque or iridescent with lumina large and transparent. Spore surface foveolate.
On tree trunks or rocks in forests, in limestone crevices; sea level to 1000 m. Anhui, Chongqing, Guangdong, Guangxi, Guizhou, Hunan, Jiangsu, Jiangxi, Sichuan, Taiwan, Xinjiang, Xizang, Yunnan, Zhejiang [Japan, Korea, Vietnam].
This is a very difficult species to define, and many specimens have been misidentified as Lepisorus thunbergianus. We found that L. tosaensis can be distinguished from L. thunbergianus by the short rhizome, iridescent young scales, and much larger transparent lumina of the leaf scales. This species is always distributed at elevations less than 1000 m, while L. thunbergianus is found at relatively higher elevations.Lepisorus paohuashanensis differs from L. tosaensis by the much smaller fronds, 3-7 × 0.4-0.6 cm, but this is just an extreme form within variable populations.
别名：阔叶瓦苇;拟瓦苇;七星剑;石韦;拟瓦韦;星子蕨;
科名：水龙骨科 Polypodiaceae
属名：瓦韦属 Lepisorus
8.阔叶瓦韦（中国高等植物图鉴）拟瓦韦（台湾植物志）图版10：1-2
Lepisorus tosaensis (Makino) H. Ito in J. Jap. Bot. 11: 93. 1935; Pic Ser., Ind. Fil. Sup. 4: 177. 1965; Tagawa, Col. Illustr. Jap. Pterid. 158. Pl. 67-363. 1959; Y. L. Zhang et al., Sporae Pterid. Sin. 493. 1976; De Vol et C. M. Kuo in H. L. Li et al., Fl. Taiwan 1: 188. 1975 et Fl. Taiwan 2nd ed. 1: 493. 1994; Nakaike, Illustr. Pterid. Jap. 2: 444. f. 445. 1981; Fl. Fujian 1: 230. 1982; 陈仁钧, 安徽植物志1: 189. 图195. 1985; S. X. Xu in j, F. Cheng et G. F. Chu, Fl. Jiangxi 1: 313. 1993; Z. F. Zhang et S. Y. Zhang, Fl. Zhejiang 1: 317. f. 334. 1992; S. L. Yu et al. in Bull. Bot. Res. 16 (1): 10. 1996. ——Polypodium tosaense Makino in Bot. Mag. Tokyo 27: 127. 1913. ——Polypodium lineare Thunb. var. caudatum Makino in Bot. Mag. Tokyo 17: 78. 1903. ——Polypodium infraplanicostale Hayata, Icon. Pl. Form. 5: 315. f. 128. 1915. ——Lepisorus infraplanicostalis (Hayata) Ching in Bull. Fan Mem. Inst. Biol. Bot. 4: 92. 1933. ——Polypodium morii Hayata, Pl. Form. 8: 153. f. 83, 84. 1919. ——Polypodium haozanense Hayata, Fl. Form. 8: 152. Pl. 81, 82. 1919.
植株高约15-30厘米。根状茎短促横卧，密被卵状披针形鳞片；鳞片深棕色，大部分不透明，仅边缘有1-2行淡棕色透明的细胞。叶簇生或近生；叶柄长1-5厘米，禾秆色；叶片披针形，中部为最宽约1-2厘米，向两端渐变狭，顶端渐尖头，基部渐狭并下延，长 (10) 13-20厘米，干后淡棕色或灰绿色，革质，两面光滑无毛。主脉上下均隆起，小脉不见。孢子囊群圆形，位于主脉与叶缘之间，聚生于叶片上半部，幼时被淡棕色圆形的隔丝覆盖。
产浙江 (临安、鄞县、开化、丽水、遂昌、庆云、平阳、宁波、杭州和苍南) 、江西 (遂川、宜丰、修水、寻乌) 、安徽 (黟县) , 福建 (建德、德化、南平、崇安) 、台湾 (南投) 、广东 (增城南昆山、翁源至英德、乐昌、状元山、乳源、新兴、信宜、河源、惠阳) 、香港、广西 (龙胜、武鸣、大明山、平南、瑶山) 、云南 (昆明、墨江、勐海、屏边、顺宁、金平、河口、文山) 、贵州 (江口、梵净山、安顺、凯里) 、四川 (峨眉山、会里、屏山) 、重庆 (北碚) 、湖南 (黔阳、莽山、保靖) 、西藏 (墨脱) 。附生溪边林下树干或岩石上, 及石灰墙缝中, 海拔650-1700米。模式标本采自日本。
《Flora of China》 Vol. 2-3 (2013)
Lepisorus tosaensis (Makino) H. It? J. Jap. Bot. 11: 93. 1935.
阔叶瓦韦 kuo ye wa wei
Polypodium tosaense Makino, Bot. Mag. (Tokyo) 27: 127. 1913; Lepisorus infraplanicostalus (Hayata) Ching; L. paohuashanensis Ching; L. sinicus Ching & Z. Y. Liu; Polypodium hoozanense Hayata; P. infraplanicostale Hayata; P. lineare N. L. Burman var. caudatum Makino; P. morii Hayata.
Plants 15-30 cm tall. Rhizomes short and decumbent, less than 3 cm, 2-3 mm in diam., apex densely scaly, remainder soon naked; scales basifixed, deep brown with paler margins, iridescent when young, lanceolate or broadly lanceolate, 2-3.5 × 0.7-1.1 mm; most lumina opaque, marginal 1 or 2 rows of cells brownish, transparent. Fronds clustered; stipe straw-colored, 1-3(-5) cm, 1-1.5 mm in diam.; lamina brownish or grayish green when dried, lanceolate or linear-lanceolate, both ends attenuate, (10-)13-20 × 1-2 cm, widest at middle, leathery, both surfaces glabrous, base cuneate, decurrent, apex acuminate; costa raised on both sides, veinlets obscure. Sori on distal half of lamina, costular, rarely midway between costa and margins, orbicular, 3-7 mm in diam.; paraphyses orbicular, brownish with central lumina small and opaque or iridescent with lumina large and transparent. Spore surface foveolate.
On tree trunks or rocks in forests, in limestone crevices; sea level to 1000 m. Anhui, Chongqing, Guangdong, Guangxi, Guizhou, Hunan, Jiangsu, Jiangxi, Sichuan, Taiwan, Xinjiang, Xizang, Yunnan, Zhejiang [Japan, Korea, Vietnam].
This is a very difficult species to define, and many specimens have been misidentified as Lepisorus thunbergianus. We found that L. tosaensis can be distinguished from L. thunbergianus by the short rhizome, iridescent young scales, and much larger transparent lumina of the leaf scales. This species is always distributed at elevations less than 1000 m, while L. thunbergianus is found at relatively higher elevations.Lepisorus paohuashanensis differs from L. tosaensis by the much smaller fronds, 3-7 × 0.4-0.6 cm, but this is just an extreme form within variable populations.