小叶海金沙Lygodium scandens (Linn.) Sw.
小叶海金沙Lygodium scandens (Linn.) Sw.
5. 小叶海金沙（中国主要植物图说，蕨类植物门）
Lygodium scandens (Linn. ) Sw. in Schrad. Journ. (1801) 106; Bedd. Ferns S. Ind. (1861) t. 61; Hook. et Bak. Syn. Fil. (1864) 437, pro parte; Prantl, Schiz. (1881) 61; Christ, Farnkr. d. Erde (1897) 354; Diels in Engl. u. Prantl,. Nat. Pflanzenfam. I, iv (1899) 366; C. Chr. Ind. Fil. (1905) 413; Dunn & Tutch. Fl. Kwangt. & Hongk. in Kew Bull. Add. Ser. X (1912) 356; Ogata, Ic. Fil. Jap. VII (1936) t. 324; Tard. -Blot et C. Chr. in Fl. Indo-Chine VII, ii (1939) 41; H. Ito, Fil. Jap. Illustr. (1944) t. 490; Holt. Fl. Mal. II, Ferns Mal. (1954) 58; 傅书遐, 中国主要植物图说 (蕨类植物门) (1957) 29页, 33图——phioglossum scandens Linn. Sp. Pl. II(1753)1063——Odontopteris scandens Bernh. in Schrad. Journ. (1801) 127——Ramondia scandens Mirbel in Bull. Soc. Philom. II (1801) 179——Hydroglossum scandens Willd. in Schr. Akad. Erfurt (1802) 20——Ugena microphylla Cav. Icon. Descr. Pl. VI (1801) 76, t. 595, f. 2——Lygodium microphyllum R. Br. Prod. Fl. N. Holl. (1810) 162; Clarke in Trans. Linn. Soc. II, Bot. I (1880) 583; Bedd. Handb. Ferns Brit. Ind. (1883) 455, f. 282.
植株蔓攀，高达5-7米。叶轴纤细如铜丝，二回羽状；羽片多数，相距约7-9厘米，羽片对生于叶轴的距上，距长2-4毫米，顶端密生红棕色毛。不育羽片生于叶轴下部，长圆形，长7-8厘米，宽4-7厘米，柄长1-1.2厘米，奇数羽状，或顶生小羽片有时两叉，小羽片4对，互生，有2-4毫米长的小柄，柄端有关节，各片相距约8毫米，卵状三角形、阔披针形或长长圆形，先端钝，基部较阔，心脏形，近平截或圆形。边缘有矮钝齿，或锯齿不甚明显。叶脉清晰，三出，小脉2-3回二叉分歧，斜向上，直达锯齿。叶薄草质，干后暗黄绿色，两面光滑。能育羽片长圆形，长8-10厘米，宽4-6厘米，通常奇数羽状，小羽片的柄长2-4毫米，柄端有关节，9-11片，互生，各片相距7-10毫米，三角形或卵状三角形，钝头，长1.5-3厘米，宽1.5-2厘米。孢子囊穗排列于叶缘，到达先端，5-8对，线形，一般长3-5毫米，最长的达8-10毫米，黄褐色，光滑。
产于福建西部（永定）、台湾（台北）、广东（惠阳、英德）、香港、海南岛西北部及南部（儋州、乐平）、广西（临桂、瑶山）、云南东南部（蒙自、河口）。产溪边灌木丛中，海拔110-152米。也分布于印度南部、缅甸、南洋羣岛、菲律宾。
《Flora of China》 Vol. 2-3 (2013)
Lygodium microphyllum (Cavanilles) R. Brown Prodr. 162. 1810.
小叶海金沙 xiao ye hai jin sha
Ugena microphylla Cavanilles, Icon. 6: 76. 1801; Lygodium scandens (Linnaeus) Swartz var. microphyllum (Cavanilles) Luerssen.
Rhizome long creeping, ca. 3 mm in diam., branching, covered with multicellular hairs 2-3 mm, lustrous black changing to reddish brown on fronds. Fronds 3-10 mm apart, climbing to 10-15 m. Stipes dark brown at base becoming stramineous, covered with reddish brown multicellular hairs, sparse distally, ca. 14 cm to first pinnae. Rachis ca. 1 mm in diam., narrowly grooved, glabrous to randomly hairy. Pinna stalk 3-6 mm, grooved, glabrous to sparsely hairy, hairs short, acicular extending onto pinna branch for a short distance. Dormant pinna bud prominent, covered with golden to reddish brown multicellular hairs. Primary pinna branches grooved, glabrous, pinnate, bearing 4(-7) alternate pairs of segments, terminal segments bifid or dichotomous. Segment stalk 2-4(-7) mm, articulate (segment deciduous leaving persistent stalk), articulation zone prominent at base of segment. Segments deltoid to lanceolate, rarely lobed, 1.5-2.5(-5.5) × 0.9-1.5(-2) cm, truncate to cuneate at base, rounded to acute at apex. Veins free, prominent abaxially, 2 or 3 times forked, ascending at 30°-40° from costa, ending at margin. Margins minutely crenulate-serrulate to subentire, slightly thickened. Fertile and sterile segments monomorphic to slightly dimorphic. Fertile segments smaller, 1-3 × 1-1.5 cm, articulation zone of stalk less prominent, hairs more common abaxially close to sorophores which extend from veins at margins, 4-6(-10) sporangial pairs per sorophore; indusia glabrous, margins crenulate; spores reticulate, laesurae not prominent.
Among shrubs by streams; low elevations (near sea level to 700 m in Taiwan). Fujian, Guangdong, Guangxi, Hainan, Taiwan, Yunnan [India, Indonesia, Malaysia, Myanmar, Nepal, Philippines; Africa, Australia, North America, S Pacific islands].
别名：small-leaf climbing fern;斑鸠窝;扫把藤;
科名：海金沙科 Lygodiaceae
属名：海金沙属 Lygodium
5. 小叶海金沙（中国主要植物图说，蕨类植物门）
Lygodium scandens (Linn. ) Sw. in Schrad. Journ. (1801) 106; Bedd. Ferns S. Ind. (1861) t. 61; Hook. et Bak. Syn. Fil. (1864) 437, pro parte; Prantl, Schiz. (1881) 61; Christ, Farnkr. d. Erde (1897) 354; Diels in Engl. u. Prantl,. Nat. Pflanzenfam. I, iv (1899) 366; C. Chr. Ind. Fil. (1905) 413; Dunn & Tutch. Fl. Kwangt. & Hongk. in Kew Bull. Add. Ser. X (1912) 356; Ogata, Ic. Fil. Jap. VII (1936) t. 324; Tard. -Blot et C. Chr. in Fl. Indo-Chine VII, ii (1939) 41; H. Ito, Fil. Jap. Illustr. (1944) t. 490; Holt. Fl. Mal. II, Ferns Mal. (1954) 58; 傅书遐, 中国主要植物图说 (蕨类植物门) (1957) 29页, 33图——phioglossum scandens Linn. Sp. Pl. II(1753)1063——Odontopteris scandens Bernh. in Schrad. Journ. (1801) 127——Ramondia scandens Mirbel in Bull. Soc. Philom. II (1801) 179——Hydroglossum scandens Willd. in Schr. Akad. Erfurt (1802) 20——Ugena microphylla Cav. Icon. Descr. Pl. VI (1801) 76, t. 595, f. 2——Lygodium microphyllum R. Br. Prod. Fl. N. Holl. (1810) 162; Clarke in Trans. Linn. Soc. II, Bot. I (1880) 583; Bedd. Handb. Ferns Brit. Ind. (1883) 455, f. 282.
植株蔓攀，高达5-7米。叶轴纤细如铜丝，二回羽状；羽片多数，相距约7-9厘米，羽片对生于叶轴的距上，距长2-4毫米，顶端密生红棕色毛。不育羽片生于叶轴下部，长圆形，长7-8厘米，宽4-7厘米，柄长1-1.2厘米，奇数羽状，或顶生小羽片有时两叉，小羽片4对，互生，有2-4毫米长的小柄，柄端有关节，各片相距约8毫米，卵状三角形、阔披针形或长长圆形，先端钝，基部较阔，心脏形，近平截或圆形。边缘有矮钝齿，或锯齿不甚明显。叶脉清晰，三出，小脉2-3回二叉分歧，斜向上，直达锯齿。叶薄草质，干后暗黄绿色，两面光滑。能育羽片长圆形，长8-10厘米，宽4-6厘米，通常奇数羽状，小羽片的柄长2-4毫米，柄端有关节，9-11片，互生，各片相距7-10毫米，三角形或卵状三角形，钝头，长1.5-3厘米，宽1.5-2厘米。孢子囊穗排列于叶缘，到达先端，5-8对，线形，一般长3-5毫米，最长的达8-10毫米，黄褐色，光滑。
产于福建西部（永定）、台湾（台北）、广东（惠阳、英德）、香港、海南岛西北部及南部（儋州、乐平）、广西（临桂、瑶山）、云南东南部（蒙自、河口）。产溪边灌木丛中，海拔110-152米。也分布于印度南部、缅甸、南洋羣岛、菲律宾。
《Flora of China》 Vol. 2-3 (2013)
Lygodium microphyllum (Cavanilles) R. Brown Prodr. 162. 1810.
小叶海金沙 xiao ye hai jin sha
Ugena microphylla Cavanilles, Icon. 6: 76. 1801; Lygodium scandens (Linnaeus) Swartz var. microphyllum (Cavanilles) Luerssen.
Rhizome long creeping, ca. 3 mm in diam., branching, covered with multicellular hairs 2-3 mm, lustrous black changing to reddish brown on fronds. Fronds 3-10 mm apart, climbing to 10-15 m. Stipes dark brown at base becoming stramineous, covered with reddish brown multicellular hairs, sparse distally, ca. 14 cm to first pinnae. Rachis ca. 1 mm in diam., narrowly grooved, glabrous to randomly hairy. Pinna stalk 3-6 mm, grooved, glabrous to sparsely hairy, hairs short, acicular extending onto pinna branch for a short distance. Dormant pinna bud prominent, covered with golden to reddish brown multicellular hairs. Primary pinna branches grooved, glabrous, pinnate, bearing 4(-7) alternate pairs of segments, terminal segments bifid or dichotomous. Segment stalk 2-4(-7) mm, articulate (segment deciduous leaving persistent stalk), articulation zone prominent at base of segment. Segments deltoid to lanceolate, rarely lobed, 1.5-2.5(-5.5) × 0.9-1.5(-2) cm, truncate to cuneate at base, rounded to acute at apex. Veins free, prominent abaxially, 2 or 3 times forked, ascending at 30°-40° from costa, ending at margin. Margins minutely crenulate-serrulate to subentire, slightly thickened. Fertile and sterile segments monomorphic to slightly dimorphic. Fertile segments smaller, 1-3 × 1-1.5 cm, articulation zone of stalk less prominent, hairs more common abaxially close to sorophores which extend from veins at margins, 4-6(-10) sporangial pairs per sorophore; indusia glabrous, margins crenulate; spores reticulate, laesurae not prominent.
Among shrubs by streams; low elevations (near sea level to 700 m in Taiwan). Fujian, Guangdong, Guangxi, Hainan, Taiwan, Yunnan [India, Indonesia, Malaysia, Myanmar, Nepal, Philippines; Africa, Australia, North America, S Pacific islands].