藤石松Lycopodiastrun Lycopodiastrum (Spring) Holub ex Dixit
藤石松Lycopodiastrun Lycopodiastrum (Spring) Holub ex Dixit
藤石松（云南植物研究） 石子藤（第尔氏华中植物），石子藤石松（中国高等植物图鉴），木贼叶石松（台湾植物志），舒筋草（四川中药志） 图版19: 1-8
Lycopodiastrum casuarinoides (Spring) Holub ex Dixit in J. Bombay Nat. Hist. Soc. 77(3): 540. 1981, et Census Ind. Pterid. 10. 1984, et Lycop. Ind. 94. f. 25B. pl. VI: f. 25. 1988; B. Olig. Index Lycop. 25. 1989; Bir et al. in Ind. Fern J. 6: 34. 1989; L. B. Zhang et H. S. Kung in Acta Phytotax. Sin. 38(3): 274. 2000. —— L. casuarinoides (Spring) Holub in Preslia 47: 103. 1975. nom nud, et in Folia Geobot. Phytotax. 18: 441. 1983; Ching in Acta Bot. Yunnan. 4(3): 225. 1982; H. S. Kung, Fl. Sichuan. 6: 54. Pl. 16: 1-4. 1988; P. S. Wang, Pterid. Fanj ing Mt. Nat. Res. 20. 1992; Fl. Dulongj iang Reg. 2. 1993; in Fl. Jiangxi 1: 23. f. 7. 1993; Fl. Zhejiang1: 9. f. 1-9. 1993: Vasc. Pl. Hengduan Mts. 1: 6. 1993. —— Lycopodium casuari-noides Spring, Monogr. Lycop. 1: 94. 1842; C. B. Clarke in Trans. Linn, Soc. II. Bot. 1: 593. 1880; Baker, Handb. Fern Allies 24. 1887; Christ in Bull. Acad. Geogr. Bot. 11: 371. 1902; Alderw, Malay. Fern-allies 49. 1915; C. Chr. in Meddel. Fran. Goteb. Bot. Tradg. 109. 1924; Nessel, Baerlappge. 371. 1939; Alston in Lecomte, Fl. Indo-Chine 7(2): 554. 1951; S. H. Fu, Ill. Important Chinese Pl., Pterid. 5. 1957; Tagawa, Col, Ill, Jap. Pterid. 15. Pl. 3: 19. 1959: Fl. Hainan. 1: 8. 1964; Ic. Corm. Sin. 1: 110, f. 219. 1972; De Vol in H. L. Li, Fl. Taiwan 1: 40, Pl. 7. 1975; Y. L. Chang et al, Sporae Pterid. Sin. 40, f.7, pl. 2: 24-25. 1976; Fl. Fukien 1: 7, f.3. 1982; Nakaike, N. Fl. Jap. Pterid. 14, f.14 a-b. 1982; P. S. Wang et X. Y. Wangin Guizhou Sci. 2: 49. 1983; C. M. Kuo in Taiwania 30: 12. 1985; B. Ollg. in OperaBot. 92: 172. 1987, et Index Lycop. 36. 1989; Fl. Fujian. rev. 1: 7, f.3. 1991: K. Iwats., Ferns Fern Allies Jap. 48, pl. 6: 4, et pl. 7: 5. 1992; J. L. Tsai et W. C.Shieh in T. C. Huang, Fl. Taiwan ed. 2, 1: 31, pl. 2. photo 34. 1994.
大型土生植物。地下茎长而匍匐。地上主茎木质藤状，伸长攀援达数米，圆柱形，直径约2毫米，具疏叶；叶螺旋状排列，贴生，卵状披针形至钻形，长1.5-3.0毫米，宽约0.5毫米，基部突出，弧形，无柄，先端渐尖，具1膜质，长2-5毫米的长芒或芒脱落。不育枝柔软，黄绿色，圆柱状，枝连叶宽约4毫米，多回不等位二叉分枝；叶螺旋状排列，但叶基扭曲使小枝呈扁平状，密生，上斜，钻状，上弯，长2-3毫米，宽约0.5毫米，基部下延，无柄，先端渐尖，具长芒，边缘全缘，背部弧形，腹部有凹槽，无光泽，中脉不明显，草质。能育枝柔软，红棕色，小枝扁平，多回二叉分枝；叶螺旋状排列，稀疏，贴生，鳞片状，长约0.8毫米，宽约0.3毫米，基部下延，无柄，先端渐尖，具芒，边缘全缘；苞片形同主茎，仅略小；孢子囊穗每6-26个一组生于多回二叉分枝的孢子枝顶端，排列成圆锥形，具直立的总柄和小柄，弯曲，长1-4厘米，直径2-3毫米，红棕色；孢子叶阔卵形，覆瓦状排列，长2-3毫米，宽约1.5毫米，先端急尖，具膜质长芒，边缘具不规则钝齿，厚膜质；孢子囊生于孢子叶腋，内藏，圆肾形，黄色。
产华东、华南、华中及西南大部分省区。生于海拔100-3100米的林下、林缘、灌丛下或沟边。亚洲其他热带及亚热带地区有分布。模式标本采自菲律宾。
《Flora of China》 Vol. 2-3 (2013)
Lycopodiastrum casuarinoides (Spring) Holub ex R. D. Dixit J. Bombay Nat. Hist. Soc. 77: 541. 1981.
藤石松 teng shi song
Lycopodium casuarinoides Spring, Monogr. Lycopod. 1: 94. 1842; Lepidotis casuarinoides (Spring) Rothmaler.
Plants large, terrestrial. Rhizome long and creeping, aerial shoots woody and vinelike, up to several meters, terete, ca. 2 mm in diam., with sparse leaves; leaves spirally arranged, adnate, ovate-lanceolate to subulate, 1.5-3 × ca. 0.5 mm, base raised, arc-shaped, sessile, apex acuminate, with 1 membranous long awn 2-5 mm, sometimes awns deciduous. Sterile branches soft, yellowish green, terete, together with leaves ca. 4 mm wide, dichotomously branched with well-differentiated lateral branchlets much branched; leaves spirally arranged, but leaf base twisted and making branchlets complanate, dense, ascending, not lustrous, subulate, angled upward, 2-3 × ca. 0.5 mm, abaxially arc-shaped, concave adaxially, midrib indistinct, herbaceous, base decurrent, sessile, margin entire, apex acuminate, long aristate. Fertile branches soft, reddish brown, branchlets flattened, dichotomously branched with well-differentiated lateral branchlets; leaves spirally arranged, sparse, appressed, scalelike, ca. 0.8 × 0.3 mm, base decurrent, sessile, margin entire, apex acuminate, aristate; bracts similar to leaves on aerial shoots but smaller; strobili 6-26 per peduncle and terminal on multi-dichotomously branched peduncles forming a cone shape, with erect peduncles and pedicels, bent, reddish brown, 1-4 cm × 2-3 mm; sporophylls broadly ovate, imbricate, 2-3 × ca. 1.5 mm, thickly membranous, with irregular teeth on margin, apex acute, with membranous long awn. Sporangia borne in axils of sporophylls, enclosed, yellow, reniform.
Forests, forest margins, shrubs, riverbanks; 100-3100 m. Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang [tropical and subtropical regions of Asia: Bhutan, India, Japan, Nepal, and SE Asia extending to Papua New Guinea].
别名：小伸筋草;天梭罗;猫公藤;木贼叶石松;舒筋草;石子藤;灯笼草;石子藤石松;藤子石松;吊壁伸筋;无病苹;伸筋草;
科名：石松科 Lycopodiaceae
属名：藤石松属 Lycopodiastrum
藤石松（云南植物研究） 石子藤（第尔氏华中植物），石子藤石松（中国高等植物图鉴），木贼叶石松（台湾植物志），舒筋草（四川中药志） 图版19: 1-8
Lycopodiastrum casuarinoides (Spring) Holub ex Dixit in J. Bombay Nat. Hist. Soc. 77(3): 540. 1981, et Census Ind. Pterid. 10. 1984, et Lycop. Ind. 94. f. 25B. pl. VI: f. 25. 1988; B. Olig. Index Lycop. 25. 1989; Bir et al. in Ind. Fern J. 6: 34. 1989; L. B. Zhang et H. S. Kung in Acta Phytotax. Sin. 38(3): 274. 2000. —— L. casuarinoides (Spring) Holub in Preslia 47: 103. 1975. nom nud, et in Folia Geobot. Phytotax. 18: 441. 1983; Ching in Acta Bot. Yunnan. 4(3): 225. 1982; H. S. Kung, Fl. Sichuan. 6: 54. Pl. 16: 1-4. 1988; P. S. Wang, Pterid. Fanj ing Mt. Nat. Res. 20. 1992; Fl. Dulongj iang Reg. 2. 1993; in Fl. Jiangxi 1: 23. f. 7. 1993; Fl. Zhejiang1: 9. f. 1-9. 1993: Vasc. Pl. Hengduan Mts. 1: 6. 1993. —— Lycopodium casuari-noides Spring, Monogr. Lycop. 1: 94. 1842; C. B. Clarke in Trans. Linn, Soc. II. Bot. 1: 593. 1880; Baker, Handb. Fern Allies 24. 1887; Christ in Bull. Acad. Geogr. Bot. 11: 371. 1902; Alderw, Malay. Fern-allies 49. 1915; C. Chr. in Meddel. Fran. Goteb. Bot. Tradg. 109. 1924; Nessel, Baerlappge. 371. 1939; Alston in Lecomte, Fl. Indo-Chine 7(2): 554. 1951; S. H. Fu, Ill. Important Chinese Pl., Pterid. 5. 1957; Tagawa, Col, Ill, Jap. Pterid. 15. Pl. 3: 19. 1959: Fl. Hainan. 1: 8. 1964; Ic. Corm. Sin. 1: 110, f. 219. 1972; De Vol in H. L. Li, Fl. Taiwan 1: 40, Pl. 7. 1975; Y. L. Chang et al, Sporae Pterid. Sin. 40, f.7, pl. 2: 24-25. 1976; Fl. Fukien 1: 7, f.3. 1982; Nakaike, N. Fl. Jap. Pterid. 14, f.14 a-b. 1982; P. S. Wang et X. Y. Wangin Guizhou Sci. 2: 49. 1983; C. M. Kuo in Taiwania 30: 12. 1985; B. Ollg. in OperaBot. 92: 172. 1987, et Index Lycop. 36. 1989; Fl. Fujian. rev. 1: 7, f.3. 1991: K. Iwats., Ferns Fern Allies Jap. 48, pl. 6: 4, et pl. 7: 5. 1992; J. L. Tsai et W. C.Shieh in T. C. Huang, Fl. Taiwan ed. 2, 1: 31, pl. 2. photo 34. 1994.
大型土生植物。地下茎长而匍匐。地上主茎木质藤状，伸长攀援达数米，圆柱形，直径约2毫米，具疏叶；叶螺旋状排列，贴生，卵状披针形至钻形，长1.5-3.0毫米，宽约0.5毫米，基部突出，弧形，无柄，先端渐尖，具1膜质，长2-5毫米的长芒或芒脱落。不育枝柔软，黄绿色，圆柱状，枝连叶宽约4毫米，多回不等位二叉分枝；叶螺旋状排列，但叶基扭曲使小枝呈扁平状，密生，上斜，钻状，上弯，长2-3毫米，宽约0.5毫米，基部下延，无柄，先端渐尖，具长芒，边缘全缘，背部弧形，腹部有凹槽，无光泽，中脉不明显，草质。能育枝柔软，红棕色，小枝扁平，多回二叉分枝；叶螺旋状排列，稀疏，贴生，鳞片状，长约0.8毫米，宽约0.3毫米，基部下延，无柄，先端渐尖，具芒，边缘全缘；苞片形同主茎，仅略小；孢子囊穗每6-26个一组生于多回二叉分枝的孢子枝顶端，排列成圆锥形，具直立的总柄和小柄，弯曲，长1-4厘米，直径2-3毫米，红棕色；孢子叶阔卵形，覆瓦状排列，长2-3毫米，宽约1.5毫米，先端急尖，具膜质长芒，边缘具不规则钝齿，厚膜质；孢子囊生于孢子叶腋，内藏，圆肾形，黄色。
产华东、华南、华中及西南大部分省区。生于海拔100-3100米的林下、林缘、灌丛下或沟边。亚洲其他热带及亚热带地区有分布。模式标本采自菲律宾。
《Flora of China》 Vol. 2-3 (2013)
Lycopodiastrum casuarinoides (Spring) Holub ex R. D. Dixit J. Bombay Nat. Hist. Soc. 77: 541. 1981.
藤石松 teng shi song
Lycopodium casuarinoides Spring, Monogr. Lycopod. 1: 94. 1842; Lepidotis casuarinoides (Spring) Rothmaler.
Plants large, terrestrial. Rhizome long and creeping, aerial shoots woody and vinelike, up to several meters, terete, ca. 2 mm in diam., with sparse leaves; leaves spirally arranged, adnate, ovate-lanceolate to subulate, 1.5-3 × ca. 0.5 mm, base raised, arc-shaped, sessile, apex acuminate, with 1 membranous long awn 2-5 mm, sometimes awns deciduous. Sterile branches soft, yellowish green, terete, together with leaves ca. 4 mm wide, dichotomously branched with well-differentiated lateral branchlets much branched; leaves spirally arranged, but leaf base twisted and making branchlets complanate, dense, ascending, not lustrous, subulate, angled upward, 2-3 × ca. 0.5 mm, abaxially arc-shaped, concave adaxially, midrib indistinct, herbaceous, base decurrent, sessile, margin entire, apex acuminate, long aristate. Fertile branches soft, reddish brown, branchlets flattened, dichotomously branched with well-differentiated lateral branchlets; leaves spirally arranged, sparse, appressed, scalelike, ca. 0.8 × 0.3 mm, base decurrent, sessile, margin entire, apex acuminate, aristate; bracts similar to leaves on aerial shoots but smaller; strobili 6-26 per peduncle and terminal on multi-dichotomously branched peduncles forming a cone shape, with erect peduncles and pedicels, bent, reddish brown, 1-4 cm × 2-3 mm; sporophylls broadly ovate, imbricate, 2-3 × ca. 1.5 mm, thickly membranous, with irregular teeth on margin, apex acute, with membranous long awn. Sporangia borne in axils of sporophylls, enclosed, yellow, reniform.
Forests, forest margins, shrubs, riverbanks; 100-3100 m. Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang [tropical and subtropical regions of Asia: Bhutan, India, Japan, Nepal, and SE Asia extending to Papua New Guinea].