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灰背老鹳草Geranium wlassowianum Fisch. ex Link

灰背老鹳草Geranium wlassowianum Fisch. ex Link

别名:绒背老鹳草;灰背老观草;柴布日-西木德格来;

科名:牻牛儿苗科 Geraniaceae

属名:老鹳草属 Geranium

《中国植物志》第43(1)卷077页
  51. 灰背老鹳草(中国高等植物图鉴)图版23: 6-12
  Geranium wlassowianum Fisch. ex Link. Enum. Hort. Berol. 2, 197. 1822; Kom Fl. Mansh. 2(2): 649. 178. 1904; R. Knuth in Engl. Pflanzenr. Heft 53. 4 (129): 178. 1912, Bobr. in Schischk. et Bobr. Fl. URSS 14: 41. 1949; 中国高等植物图鉴2: 528. 图2785. 1972; 东北草本植物志6: 14. 1979; 内蒙古植物志4: 8. 1979; Kitag. Lineam Fl. Mansh. 420. 1979.
  多年生草本,高30-70厘米。根茎短粗,木质化,斜生或直生,具簇生纺锤形块根。上部围以残存基生托叶和叶柄,茎2-3,直立或基部仰卧,具棱角,假二叉状分枝,被倒向短柔毛。叶基生和茎上对生;托叶三角状披针形或卵状披针形,长7-8毫米,宽3-4毫米,先端具芒状长尖头;基生叶具长柄,柄长为叶片的4-5倍,被短柔毛,近叶片处被毛密集,茎下部叶柄稍长于叶片,上部叶柄明显短于叶片;叶片五角状肾圆形,基部浅心形,长4-6厘米,宽6-9厘米,5深裂达中部或稍过之,裂片倒卵状楔形,下部全缘,上部3深裂,中间小裂片狭长,3裂,侧小裂片具1-3牙齿,表面被短伏毛,背面灰白色,沿脉被短糙毛。花序腋生和顶生,稍长于叶,总花梗被倒向短柔毛,具2花;苞片狭披针形,长6-8毫米,宽1-1.5毫米;花梗与总花梗相似,通常长为花的1.5-2倍,花期直立或弯曲,果期水平状叉开;萼片长卵形或矩圆形状椭圆形,长8-10毫米,宽3-4毫米,先端具长尖头,密被短柔毛和开展的疏散长柔毛;花瓣淡紫红色,具深紫色脉纹,宽倒卵形,长约为萼片的2倍,先端圆形,基部楔形,被长柔毛;雄蕊稍长于萼片,花丝棕褐色,下部扩展,边缘和基部被长糙毛,花药棕褐色;雌蕊被短糙毛,花柱分枝棕褐色,与花柱近等长。蒴果长约3厘米,被短糙毛。花期7-8月,果期8-9月。分布于东北、山西、河北、山东和内蒙古等。生于低、中山的山地草甸、林缘等处。俄罗斯东西伯利亚、贝加尔、远东,蒙古,朝鲜皆有分布。
  本种的向南分布仅达山西,过去有关分布于湖北和四川等地的记载可能是由于鉴定不准确引起的。实际上,本种是湖北老鹳草 G. rosthornii R. Knuth的地理替代种。

《Flora of China》 Vol. 11 (2008)
Geranium wlassovianum  Fischer ex Link
灰背老鹳草   hui bei lao guan cao
Perennials. Rootstock ± vertical, 0.7-1.7 cm in diam., not tuberculate, with a fascicle of long thickened roots. Stem 25-86 cm tall, erect, not rooting at nodes, with 0.3-0.9 mm patent or rarely retrorse and appressed nonglandular trichomes. Stipules lanceolate, distinct or connate at least at base. Leaves opposite; petiole with usually 0.2-0.9 mm patent nonglandular trichomes; leaf blade 3.7-8.1 cm, palmately cleft, ratio of main sinus/middle segment length = 0.7-0.81(-0.84), pilose with appressed nonglandular trichomes; segments 5, rhombic, 6-13(-19.1) mm wide at base, 7-10-lobed in distal half, ratio of second sinus/middle segment length = (0.13-)0.2-0.24(-0.25). Cymules solitary, 2-flowered; peduncle 4.3-12.3 cm. Pedicel 2-6.2 cm, with 0.2-0.6 mm patent or retrorse ± appressed nonglandular trichomes; bracteoles lanceolate. Sepals (0.8-)0.9-1.1 cm, mucro 1.1-3.9 mm, ratio of mucro/sepal length = 0.1-0.3, with 0.1-0.3 mm antrorse ± appressed nonglandular trichomes and 0.2-1 mm ± patent nonglandular trichomes on abaxial side, glabrous adaxially. Petals 1.6-2.1 cm, erect to patent, entire, adaxial base with trichomes, margin basally ciliate, deep purplish magenta. Staminal filaments pinkish, lanceolate, base slightly dilated, abaxially densely pilose and proximal half ciliate, trichomes 0.6-1.5 mm; anthers bluish, 1.6-2.6 mm. Nectaries 5, hemispheric, abaxially glabrous, apex with a tuft of trichomes. Stigma deep red to pink. Fruit 3-3.4 cm, erect when immature; mericarps smooth, with a basal callus, with 0.3-0.7 mm ± patent nonglandular trichomes and apically usually with 0.6-1 mm patent glandular trichomes; rostrum 2.1-2.5 cm, with a 3-4.8 mm narrowed apex; stigmatic remains 5-6.9 mm. Seeds 2-2.5 mm. Fl. Jul-Aug, fr. Aug-Sep. 2n = 28, 56.
Forests, meadows; 1800-3400 m. Hebei, Heilongjiang, Henan, Jilin, Liaoning, Nei Mongol, Shandong, Shanxi [Korea, Mongolia, Russia (Baikal to Far East)].
Geranium wlassovianum and G. maximowiczii share shallowly divided leaves, paler on the abaxial side. Both species are difficult to differentiate, and the main characters are referred to in the key. Additionally, in G. wlassovianum, the anthers, gynoecium, fruit, and staminal filaments are longer (and with a dense tuft of trichomes). Trichomes on the stem and inflorescence are also longer and usually patent.

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