菊状千里光Senecio laetus Edgew.
菊状千里光Senecio laetus Edgew.
43.菊状千里光 图版62: 1-5
Senecio laetus Edgew. in Trans, Linn. Soc. Bot. 20: 74. 1846; C. Jeffrey et Y. L. Chen in Kew Bull. 39 (2): 404. 1984; 贵州植物志9: 253.图版59: 1-4. 1989; 横断山区维管植物下册: 2100. 1994. ——Senecio chrysanthemoides DC. Prodr. 6: 365. 1838; nom. illegit., non Schrank 1789; Hook. f. Fl. Brit. Ind. 3: 339. 1881; Ling in Contr. Inst. Bot. Nat. Acad. Peiping. 3: 127. 1935; Hand. -Maaz. Symb. Sin. 7. 1120. 1936; Kitam. in Hara Fl. East. Himal. 343. 1966; S. Y. Hu in Quart. Journ. Taiwan Mus. 21: 40. 1968; Koyama in Mem. Fac. Sc. Kyoto Univ. Ser. Biol. 2 (2): 154. 1969; 中国高等植物图鉴4: 571.图6556, 1975; Kitam. in Hara Fl. East. Himal. 2: 140. 1971, 3: 118. 1975; Kitam. et Gould in Hara, Enumer. Flower. Pl. Nepal 3: 41. 1982. ——Senecio chrysanthemoides DC. var. khasianus (C. B. Clarke) Hook. f. Fl. Brit. Ind. 3: 339. 1881; Diels in Bot. Jahrb 29: 620. 1901; S. Y. Hu in Quart. Jorun. Taiwan Mus. 21: 40. 1968. ——Senecio argunensis auct. non Turcz.: Hand. -Mazz. Symb. Sin. 7: 1120. 1936, ——Senecio chrysanthemoides DC. var. eustegius Hand. -Mazz. in Notizbl. Bot. Gart. Berlin 13: 639. 1937; S. Y. Hu in Quart. Journ. Taiwan Mus. 21: 40. 1968. ——Senecio rosulifer Levl. in Repert Sp. Nov. 8: 359. 1910, p. p., quoad Cavalerie 3701 solum. ——Senecio scapiformis Y. L. Chen et K. Y. Pan in Acta Phytotax. Sin. 19: 93. 1981; 西藏植物志4: 818. 1985.
多年生根状茎草本，具茎叶，稀近葶状。茎单生，直立，高40-80厘米，不分枝或有花序枝，被疏蛛丝状毛，或变无毛。基生叶在花期生存或凋落。基生叶和最下部茎叶具柄，全形卵状椭圆形，卵状披针形至倒披针形，长8-10（-20）厘米，宽3-7厘米，顶端钝，基部微心形至楔状狭，具齿，不分裂或大头羽状分裂，侧裂片1-4对，向叶基部小，纸质，上面无毛，下面有疏蛛丝状毛，或多少变无毛，羽状脉，侧脉8-9对；叶柄长达10厘米，基部扩大；中部茎叶全形长圆形或倒披针状长圆形，长5-22厘米，宽2-7厘米，大头羽状浅裂或羽状浅裂，裂片多变异，顶生裂片大至小，卵形至长圆状披针形，具齿或细裂，侧裂片5-8对，长圆形至狭长圆状披针形，全缘或通常有不规则锯齿状齿或细裂，开展或稍上升，基部具耳；耳具齿或细裂，半抱茎；上部叶渐小，长圆状披针形或长圆状线形，具粗羽状齿。头状花序有舌状花，多数，排列成顶生伞房花序或复伞房花序；花序梗长5-25毫米，或多或少被蛛丝状绒毛或黄褐色短柔毛，或变无毛，有线形苞片和2-3线状钻形小苞片。总苞钟状，长3-4毫米，宽3-4毫米，具外层苞片；苞片8-10，线状钻形，长2-3毫米；总苞片10-13，长圆状披针形，渐尖，上端黑褐色，有柔毛，草质，边缘宽干膜质，向基部有黄褐色短柔毛。舌状花10-13，管部长2.5毫米；舌片黄色，长圆形，长约6.5毫米，宽2毫米，钝，上端具3细齿，有4脉；管状花多数，花冠黄色，长5-5.5毫米，管部长2.5毫米，檐部漏斗状；裂片卵状三角形，长0.8毫米，尖；花药长2毫米，基部具钝耳；附片卵状披针形；花药颈部稍伸长，向基部稍膨大。花柱分枝长1毫米，顶端截形，有乳头状毛。瘦果圆柱形，全部或管状花的瘦果有疏柔毛，有时舌状花的或全部小花的瘦果无毛。冠毛长约4毫米，污白色，禾杆色或稀淡红色，冠毛在全部瘦果存在，或有时舌状花的瘦果无冠毛；或脱落。花期4-11月。
产西藏（珠穆朗玛峰北坡、聂拉木）、重庆（綦江）、贵州（平坝、罗福、安顺、清镇、遵义、威宁、纳拥）、湖北（宜昌）、湖南（洞庭湖附近）、云南（腾冲、澜沧江一怒江分水岭、维西、丽江、澜沧、中甸、大理、顺宁、思茅、昆明、巧家、东川、蒙自、屏边）。生于林下、林缘、开旷草坡、田边和路边，海拔1 100-3750米。巴基斯坦、印度、尼泊尔和不丹也有。模式采自印度库盟。
本种自巴基斯坦西北部、印度西北和东北部至中国西南部及中部有广泛的分布。体态、叶形、瘦果的毛被、冠毛的颜色等方面多变异。舌状花瘦果具少数淡红色冠毛或无冠毛，与莱菔叶千里光S. raphanifolius Wall. ex DC.往往难以区别。此两种居群间的关系有待野外观察和研究。
《Flora of China》 Vol. 20-21 (2011)
44. Senecio analogus Candolle, Prodr. 6: 366. 1838.
菊状千里光 ju zhuang qian li guang Jacobaea analoga (Candolle) Veldkamp; Senecio chrysanthemoides Candolle (1838), not Schrank (1789); S. chrysanthemoides var. eustegius Handel-Mazzetti; S. chrysanthemoides var. khasianus (C. B. Clarke) J. D. Hooker; S. chrysanthemoides var. sisymbriiformis (Candolle) J. D. Hooker; S. chrysanthemoides var. spectabilis (Wallich ex Candolle) J. D. Hooker; S. laetus Edgeworth; S. laetus var. analogus (Candolle) R. Mathur ex Karthikeyan & Moorthy; S. laetus var. sisymbriiformis (Candolle) Aswal; S. laetus var. spectabilis (Wallich ex Candolle) R. Mathur ex Karthikeyan & Moorthy; S. pallens Wallich ex Candolle var. khasianus C. B. Clarke; S. scapiformis Y. L. Chen & K. Y. Pan; S. sisymbriiformis Candolle; S. spectabilis Wallich ex Candolle.
Herbs, perennial, rhizomatous, with leafy stems, rarely subscapigerous. Stem solitary, erect, 40-80 cm tall, simple or branching into synflorescence, sparsely arachnoid, glabrescent. Basal leaves sometimes present at or sometimes withered and shriveled or shed by anthesis; basal and lowest stem leaves petiolate; petiole to 10 cm; blade ovate-elliptic or ovate-lanceolate to oblanceolate, 8-10(-20) × 3-7 cm, papery, abaxially sparsely arachnoid and ± glabrescent, adaxially glabrescent to glabrous, pinnately veined, lateral veins 16-18, base expanded, weakly cordate to cuneate or attenuate, margin dentate, apex obtuse, unlobed or lyrate-pinnately lobed with 2-8 basal lateral lobes decreasing in size toward base of leaf. Median stem leaves oblong or oblanceolate-oblong, 5-22 × 2-7 cm, base auriculate; auricle dentate or lobulate, subamplexicaul, lyrate-pinnatifid or pinnatifid, very variable in dissection, with large to small, ovate to oblong-lanceolate, dentate or lobulate terminal lobe and 10-16 oblong to narrowly oblong-lanceolate, entire or usually irregularly serrate-dentate or lobulate spreading or somewhat ascending lateral lobes. Upper leaves smaller, oblong-lanceolate to oblong-linear, margin coarsely pinnate-dentate. Capitula radiate, numerous, arranged in terminal corymbs or compound corymbs; peduncles 5-25 mm, ± arachnoid-tomentose or fulvous puberulent, ± glabrescent, linear-bracteate, with 2 or 3 linear-subulate bracteoles. Involucres campanulate, 3-4 × 3-4 mm, calyculate; bracts of calyculus 8-10, linear-subulate, 2-3 mm; phyllaries 10-13, oblong-lanceolate, herbaceous, fulvous puberulent toward base, margin broadly scarious, apically acute or acuminate, brownish black and pubescent. Ray florets 10-13; corolla tube ca. 2.5 mm; lamina yellow, oblong, ca. 6.5 × 2 mm, 4-veined, apically obtuse, 3-denticulate. Disk florets many; corolla yellow, 5-5.5 mm, with ca. 2.5 mm tube and funnelform limb; lobes ovate-triangular, ca. 0.8 mm, apically acute. Anthers ca. 2 mm, basally obtuse-auriculate, appendages ovate-lanceolate; antheropodia slightly elongated, basally somewhat dilated. Style branches ca. 1 mm. Achenes cylindric, 2-2.5 mm, all or disk achenes sparsely pubescent, sometimes ray achenes or rarely all achenes glabrous. Pappus dirty white, straw-colored, or rarely rufous, ca. 4 mm, present in all achenes or sometimes scant or absent in ray achenes, those in ray achenes deciduous. Fl. Apr.Forests, forest and thicket margins, open grassy places, field margins, roadsides; 1100-3800 m. Guizhou, Hubei, Sichuan, Xizang, Yunnan [Bhutan, NE and NW India, Nepal, NW Pakistan].
别名：菊叶千里光;青菜一棵蒿;紫背草;野苦菜;山青菜;滇败酱;野青菜;
科名：菊科 Compositae
属名：千里光属 Senecio
43.菊状千里光 图版62: 1-5
Senecio laetus Edgew. in Trans, Linn. Soc. Bot. 20: 74. 1846; C. Jeffrey et Y. L. Chen in Kew Bull. 39 (2): 404. 1984; 贵州植物志9: 253.图版59: 1-4. 1989; 横断山区维管植物下册: 2100. 1994. ——Senecio chrysanthemoides DC. Prodr. 6: 365. 1838; nom. illegit., non Schrank 1789; Hook. f. Fl. Brit. Ind. 3: 339. 1881; Ling in Contr. Inst. Bot. Nat. Acad. Peiping. 3: 127. 1935; Hand. -Maaz. Symb. Sin. 7. 1120. 1936; Kitam. in Hara Fl. East. Himal. 343. 1966; S. Y. Hu in Quart. Journ. Taiwan Mus. 21: 40. 1968; Koyama in Mem. Fac. Sc. Kyoto Univ. Ser. Biol. 2 (2): 154. 1969; 中国高等植物图鉴4: 571.图6556, 1975; Kitam. in Hara Fl. East. Himal. 2: 140. 1971, 3: 118. 1975; Kitam. et Gould in Hara, Enumer. Flower. Pl. Nepal 3: 41. 1982. ——Senecio chrysanthemoides DC. var. khasianus (C. B. Clarke) Hook. f. Fl. Brit. Ind. 3: 339. 1881; Diels in Bot. Jahrb 29: 620. 1901; S. Y. Hu in Quart. Jorun. Taiwan Mus. 21: 40. 1968. ——Senecio argunensis auct. non Turcz.: Hand. -Mazz. Symb. Sin. 7: 1120. 1936, ——Senecio chrysanthemoides DC. var. eustegius Hand. -Mazz. in Notizbl. Bot. Gart. Berlin 13: 639. 1937; S. Y. Hu in Quart. Journ. Taiwan Mus. 21: 40. 1968. ——Senecio rosulifer Levl. in Repert Sp. Nov. 8: 359. 1910, p. p., quoad Cavalerie 3701 solum. ——Senecio scapiformis Y. L. Chen et K. Y. Pan in Acta Phytotax. Sin. 19: 93. 1981; 西藏植物志4: 818. 1985.
多年生根状茎草本，具茎叶，稀近葶状。茎单生，直立，高40-80厘米，不分枝或有花序枝，被疏蛛丝状毛，或变无毛。基生叶在花期生存或凋落。基生叶和最下部茎叶具柄，全形卵状椭圆形，卵状披针形至倒披针形，长8-10（-20）厘米，宽3-7厘米，顶端钝，基部微心形至楔状狭，具齿，不分裂或大头羽状分裂，侧裂片1-4对，向叶基部小，纸质，上面无毛，下面有疏蛛丝状毛，或多少变无毛，羽状脉，侧脉8-9对；叶柄长达10厘米，基部扩大；中部茎叶全形长圆形或倒披针状长圆形，长5-22厘米，宽2-7厘米，大头羽状浅裂或羽状浅裂，裂片多变异，顶生裂片大至小，卵形至长圆状披针形，具齿或细裂，侧裂片5-8对，长圆形至狭长圆状披针形，全缘或通常有不规则锯齿状齿或细裂，开展或稍上升，基部具耳；耳具齿或细裂，半抱茎；上部叶渐小，长圆状披针形或长圆状线形，具粗羽状齿。头状花序有舌状花，多数，排列成顶生伞房花序或复伞房花序；花序梗长5-25毫米，或多或少被蛛丝状绒毛或黄褐色短柔毛，或变无毛，有线形苞片和2-3线状钻形小苞片。总苞钟状，长3-4毫米，宽3-4毫米，具外层苞片；苞片8-10，线状钻形，长2-3毫米；总苞片10-13，长圆状披针形，渐尖，上端黑褐色，有柔毛，草质，边缘宽干膜质，向基部有黄褐色短柔毛。舌状花10-13，管部长2.5毫米；舌片黄色，长圆形，长约6.5毫米，宽2毫米，钝，上端具3细齿，有4脉；管状花多数，花冠黄色，长5-5.5毫米，管部长2.5毫米，檐部漏斗状；裂片卵状三角形，长0.8毫米，尖；花药长2毫米，基部具钝耳；附片卵状披针形；花药颈部稍伸长，向基部稍膨大。花柱分枝长1毫米，顶端截形，有乳头状毛。瘦果圆柱形，全部或管状花的瘦果有疏柔毛，有时舌状花的或全部小花的瘦果无毛。冠毛长约4毫米，污白色，禾杆色或稀淡红色，冠毛在全部瘦果存在，或有时舌状花的瘦果无冠毛；或脱落。花期4-11月。
产西藏（珠穆朗玛峰北坡、聂拉木）、重庆（綦江）、贵州（平坝、罗福、安顺、清镇、遵义、威宁、纳拥）、湖北（宜昌）、湖南（洞庭湖附近）、云南（腾冲、澜沧江一怒江分水岭、维西、丽江、澜沧、中甸、大理、顺宁、思茅、昆明、巧家、东川、蒙自、屏边）。生于林下、林缘、开旷草坡、田边和路边，海拔1 100-3750米。巴基斯坦、印度、尼泊尔和不丹也有。模式采自印度库盟。
本种自巴基斯坦西北部、印度西北和东北部至中国西南部及中部有广泛的分布。体态、叶形、瘦果的毛被、冠毛的颜色等方面多变异。舌状花瘦果具少数淡红色冠毛或无冠毛，与莱菔叶千里光S. raphanifolius Wall. ex DC.往往难以区别。此两种居群间的关系有待野外观察和研究。
《Flora of China》 Vol. 20-21 (2011)
44. Senecio analogus Candolle, Prodr. 6: 366. 1838.
菊状千里光 ju zhuang qian li guang Jacobaea analoga (Candolle) Veldkamp; Senecio chrysanthemoides Candolle (1838), not Schrank (1789); S. chrysanthemoides var. eustegius Handel-Mazzetti; S. chrysanthemoides var. khasianus (C. B. Clarke) J. D. Hooker; S. chrysanthemoides var. sisymbriiformis (Candolle) J. D. Hooker; S. chrysanthemoides var. spectabilis (Wallich ex Candolle) J. D. Hooker; S. laetus Edgeworth; S. laetus var. analogus (Candolle) R. Mathur ex Karthikeyan & Moorthy; S. laetus var. sisymbriiformis (Candolle) Aswal; S. laetus var. spectabilis (Wallich ex Candolle) R. Mathur ex Karthikeyan & Moorthy; S. pallens Wallich ex Candolle var. khasianus C. B. Clarke; S. scapiformis Y. L. Chen & K. Y. Pan; S. sisymbriiformis Candolle; S. spectabilis Wallich ex Candolle.
Herbs, perennial, rhizomatous, with leafy stems, rarely subscapigerous. Stem solitary, erect, 40-80 cm tall, simple or branching into synflorescence, sparsely arachnoid, glabrescent. Basal leaves sometimes present at or sometimes withered and shriveled or shed by anthesis; basal and lowest stem leaves petiolate; petiole to 10 cm; blade ovate-elliptic or ovate-lanceolate to oblanceolate, 8-10(-20) × 3-7 cm, papery, abaxially sparsely arachnoid and ± glabrescent, adaxially glabrescent to glabrous, pinnately veined, lateral veins 16-18, base expanded, weakly cordate to cuneate or attenuate, margin dentate, apex obtuse, unlobed or lyrate-pinnately lobed with 2-8 basal lateral lobes decreasing in size toward base of leaf. Median stem leaves oblong or oblanceolate-oblong, 5-22 × 2-7 cm, base auriculate; auricle dentate or lobulate, subamplexicaul, lyrate-pinnatifid or pinnatifid, very variable in dissection, with large to small, ovate to oblong-lanceolate, dentate or lobulate terminal lobe and 10-16 oblong to narrowly oblong-lanceolate, entire or usually irregularly serrate-dentate or lobulate spreading or somewhat ascending lateral lobes. Upper leaves smaller, oblong-lanceolate to oblong-linear, margin coarsely pinnate-dentate. Capitula radiate, numerous, arranged in terminal corymbs or compound corymbs; peduncles 5-25 mm, ± arachnoid-tomentose or fulvous puberulent, ± glabrescent, linear-bracteate, with 2 or 3 linear-subulate bracteoles. Involucres campanulate, 3-4 × 3-4 mm, calyculate; bracts of calyculus 8-10, linear-subulate, 2-3 mm; phyllaries 10-13, oblong-lanceolate, herbaceous, fulvous puberulent toward base, margin broadly scarious, apically acute or acuminate, brownish black and pubescent. Ray florets 10-13; corolla tube ca. 2.5 mm; lamina yellow, oblong, ca. 6.5 × 2 mm, 4-veined, apically obtuse, 3-denticulate. Disk florets many; corolla yellow, 5-5.5 mm, with ca. 2.5 mm tube and funnelform limb; lobes ovate-triangular, ca. 0.8 mm, apically acute. Anthers ca. 2 mm, basally obtuse-auriculate, appendages ovate-lanceolate; antheropodia slightly elongated, basally somewhat dilated. Style branches ca. 1 mm. Achenes cylindric, 2-2.5 mm, all or disk achenes sparsely pubescent, sometimes ray achenes or rarely all achenes glabrous. Pappus dirty white, straw-colored, or rarely rufous, ca. 4 mm, present in all achenes or sometimes scant or absent in ray achenes, those in ray achenes deciduous. Fl. Apr.Forests, forest and thicket margins, open grassy places, field margins, roadsides; 1100-3800 m. Guizhou, Hubei, Sichuan, Xizang, Yunnan [Bhutan, NE and NW India, Nepal, NW Pakistan].