厚喙菊Dubyaea hispida (D. Don) DC.
厚喙菊Dubyaea hispida (D. Don) DC.
3.厚喙菊 图版21: 3-4
Dubyaea hispida (D. Don) DC., Prodr. 7: 247. 1838. quoad nomen.; Stebbins in Mem. Torrey Bot. Club. 19: 18. 1940. p. p.; S. Y. Hu in Quart. Journ. Taiwan Mus. 19 (3-4): 236. 1966. p. p. ; Kitam. et S. Gould., Emumer. Flowering Plant Nepal 3: 25, 1982; 西藏植物志4: 954. 1985. quoad nomen.; Shih in Act. Phytotax. Sin. 31: 436. 1993.——Hieracium hispidum D. Don, Prodr. Fl. Nepal 165. 1825.
多年生草本，高20-100厘米，有极短的根状茎。茎直立，基部直径2-4毫米，上部有长或短的伞房花序状分枝，全部茎枝有条棱，被黑褐色的多细胞长节毛，上部及花序分枝上的毛稠密。基生叶及下部茎叶不裂，椭圆形或长椭圆形，长6-11厘米，宽3-5厘米，基部渐狭，有翼柄或无柄；中上部茎叶与基生叶及下部茎叶同形，但较小或披针形，无柄或有短翼柄；接花序下部的常线形；全部叶顶端急尖或渐尖，边缘有稀疏的凹齿，齿顶及齿缘有小尖头，但上部的叶通常全缘，无锯齿，两面被稠密或稀疏的黑色多细胞长节毛。头状花序下垂或歪斜，通常2-7枚在茎枝顶端排成伞房花序或聚伞花序。总苞宽钟状，长1.5厘米，宽1.5-2.5厘米；总苞片3层，外层长椭圆形，长8毫米，宽1毫米，中层披针形，长11毫米，宽1.5毫米，内层披针形或长椭圆形，长15毫米，宽2毫米；全部总苞片外面沿中脉被黑色或黑褐色多细胞长节毛，但向内层的毛较稀疏，外层顶端渐尖，中内层顶端急尖。舌状小花黄色。瘦果近纺锤形，压扁，上部黄色，下部黑色，长约7毫米，有多数(12-17条），不等粗纵肋，顶端截形。冠毛淡黄色，2层，易折断，长2厘米，微细锯齿。花果期8-10月。
分布云南西北部（丽江、贡山、德钦）、西藏（林芝、错那、察隅、墨脱）、四川西南部（木里、盐源）。生于高山林缘、林下、草甸或灌丛中，海拔3 200-4 200米。尼泊尔、锡金、不丹、印度北部及缅甸北部有分布。模式标本采自尼泊尔。
形态特征：多年生草本，高20一100厘米，有极短的根状茎。茎直立，基部直径2—4毫米，上部有长或短的伞房花序状分枝，全部茎枝有条棱，被黑褐色的多细胞长节毛，上部及花序分枝上的毛稠密。基生叶及下部茎叶不裂，椭圆形或长椭圆形，长6—11厘米，宽3—5厘米，基部渐狭，有翼柄或无柄；中上部茎叶与基生叶及下部茎叶同形，但较小或披针形，无柄或有短翼柄；接花序下部的常线形；全部叶顶端急尖或渐尖，边缘有稀疏的凹齿，齿顶及齿缘有小尖头，但上部的叶通常全缘，无锯齿，两面被稠密或稀疏的黑色多细胞长节毛。头状花序下垂或歪斜，通常2一?枚在茎枝顶端排成伞房花序或聚伞花序。总苞宽钟状，长1．5厘米，宽1．5—2．5厘米；总苞片3层，外层长椭圆形，长8毫米，宽1毫米，中层披针形，长ll毫米，宽1．5毫米，内层披针形或长椭圆形，长15毫米，宽2毫米；全部总苞片外面沿中脉被黑色或黑褐色多细胞长节毛，但向内层的毛较稀疏，外层顶端渐尖，中内层顶端急尖。舌状小花黄色。瘦果近纺锤形，压扁，上部黄色，下部黑色，长约7毫米，有多数(12—17条)，不等粗纵肋，顶端截形。冠毛淡黄色，2层，易折断，长2厘米，微细锯齿。花果期8—10月。
产地分布：分布云南西北部(丽江、贡山、德钦)、西藏(林芝、错那、察隅、墨脱)、四川西南部(木里、盐源)。生于高山林缘、林下、草甸或灌丛中，海拔3 200—4 200米。尼泊尔、 锡金、不丹、印度北部及缅甸北部有分布。模式标本采自尼泊尔。
参考文献：Dubyaea hispida (D．Don) DC．，Prodr．7：247．1838．Quoad nomen．；Stebbins in Mem．Torrey Bot．Club．19：18．1940．P．P．；S．Y．Hu in Quart．Journ．Taiwan Mus．19(3—4)：236．1966．P．P．；Kitam．Et 5．Gould．，Emumer．Flowering Plant Nepal 3：25，1982；西藏植物志4：954．1985．Quoadnomen．；Shih in Act．Phytotax． Sin．31：436．1993．——Hieracium hispidum D．Don，Prodr．F1．Nepal 165．1825．
《Flora of China》 Vol. 20-21 (2011)
Dubyaea hispida Candolle Prodr. 7: 247. 1838.
厚喙菊 hou hui ju
Hieracium hispidum D. Don, Prodr. Fl. Nepal. 165. 1825, not Forssk?l (1775); Crepis bhotanica Hutchinson; C. dubyaea (C. B. Clarke) C. Marquand & Airy Shaw; Dubyaea bhotanica (Hutchinson) C. Shih; D. lanceolata C. Shih; D. pteropoda C. Shih; Lactuca dubyaea C. B. Clarke.
Herbs 20-60 cm tall, perennial, with creeping ?rhizomes. Stem erect, branched from below middle or higher up, leafy, with long blackish stiff glandular hairs and pale curly weak hairs. Basal and lower stem leaves sessile or base attenuate into a long petiole-like portion, oblanceolate, elliptic, or narrowly elliptic, 5-18 × 1.5-6 cm, sinuate-dentate to shallowly lyrately pinnatifid, both faces with hairs or mainly abaxially, base narrow to widened and ± clasping, margin mucronulately dentate, apex acute to acuminate. Middle and upper stem leaves similar to lower stem leaves but ± lanceolate, smaller, base auriculately clasping; uppermost stem leaves linear-lanceolate, reduced in size. Synflorescence corymbiform, with 2-7 capitula. Capitula nodding, with 40-50 florets; peduncle blackish pilose. Involucre campanulate, 1.8-2 × 1-1.5 cm. Phyllaries with long dark brown to blackish stiff glandular hairs on midvein except on innermost ones; outer phyllaries rather few, narrowly lanceolate to linear, closely approaching inner ones in length, apex acute to acuminate; inner phyllaries many, lanceolate, margin apically often ± brownish ciliate to fringed. Florets yellow. Achene brown, 7-9 mm, with several ribs, apex long attenuate and paler. Pappus yellowish, 0.8-1.2 cm. Fl. and fr. Jul-Nov. 2n = 16.
Forests, forest margins, meadows, thickets; 2700-4500 m. Sichuan, Xizang, Yunnan [Bhutan, India (Sikkim), N Myanmar, Nepal].
Dubyaea hispida is here considered in the wider sense established by Stebbins (Mem. Torrey Bot. Club 19(3): 19. 1940) and taken up by X. Zhuang (Fl. Yunnan. 13: 704. 2004), who consequently included also the segregates D. lanceolata and D. pteropoda, distinguished only by minor differences in leaf shape. The underground parts are usually described as creeping rhizomes from which the stems arise; no specimen, however, has been seen where a true rhizome is preserved. The underground parts preserved in the material seen could also be interpreted as root-borne shoots.Dubyaea hispida is here considered in the wider sense established by Stebbins (Mem. Torrey Bot. Club 19(3): 19. 1940) and taken up by X. Zhuang (Fl. Yunnan. 13: 704. 2004), who consequently included also the segregates D. lanceolata and D. pteropoda, distinguished only by minor differences in leaf shape. The underground parts are usually described as creeping rhizomes from which the stems arise; no specimen, however, has been seen where a true rhizome is preserved. The underground parts preserved in the material seen could also be interpreted as root-borne shoots.
别名：不丹厚喙菊;披针叶厚喙菊;翼柄厚喙菊;厚缘菊;
科名：菊科 Compositae
属名：厚喙菊属 Dubyaea
3.厚喙菊 图版21: 3-4
Dubyaea hispida (D. Don) DC., Prodr. 7: 247. 1838. quoad nomen.; Stebbins in Mem. Torrey Bot. Club. 19: 18. 1940. p. p.; S. Y. Hu in Quart. Journ. Taiwan Mus. 19 (3-4): 236. 1966. p. p. ; Kitam. et S. Gould., Emumer. Flowering Plant Nepal 3: 25, 1982; 西藏植物志4: 954. 1985. quoad nomen.; Shih in Act. Phytotax. Sin. 31: 436. 1993.——Hieracium hispidum D. Don, Prodr. Fl. Nepal 165. 1825.
多年生草本，高20-100厘米，有极短的根状茎。茎直立，基部直径2-4毫米，上部有长或短的伞房花序状分枝，全部茎枝有条棱，被黑褐色的多细胞长节毛，上部及花序分枝上的毛稠密。基生叶及下部茎叶不裂，椭圆形或长椭圆形，长6-11厘米，宽3-5厘米，基部渐狭，有翼柄或无柄；中上部茎叶与基生叶及下部茎叶同形，但较小或披针形，无柄或有短翼柄；接花序下部的常线形；全部叶顶端急尖或渐尖，边缘有稀疏的凹齿，齿顶及齿缘有小尖头，但上部的叶通常全缘，无锯齿，两面被稠密或稀疏的黑色多细胞长节毛。头状花序下垂或歪斜，通常2-7枚在茎枝顶端排成伞房花序或聚伞花序。总苞宽钟状，长1.5厘米，宽1.5-2.5厘米；总苞片3层，外层长椭圆形，长8毫米，宽1毫米，中层披针形，长11毫米，宽1.5毫米，内层披针形或长椭圆形，长15毫米，宽2毫米；全部总苞片外面沿中脉被黑色或黑褐色多细胞长节毛，但向内层的毛较稀疏，外层顶端渐尖，中内层顶端急尖。舌状小花黄色。瘦果近纺锤形，压扁，上部黄色，下部黑色，长约7毫米，有多数(12-17条），不等粗纵肋，顶端截形。冠毛淡黄色，2层，易折断，长2厘米，微细锯齿。花果期8-10月。
分布云南西北部（丽江、贡山、德钦）、西藏（林芝、错那、察隅、墨脱）、四川西南部（木里、盐源）。生于高山林缘、林下、草甸或灌丛中，海拔3 200-4 200米。尼泊尔、锡金、不丹、印度北部及缅甸北部有分布。模式标本采自尼泊尔。
形态特征：多年生草本，高20一100厘米，有极短的根状茎。茎直立，基部直径2—4毫米，上部有长或短的伞房花序状分枝，全部茎枝有条棱，被黑褐色的多细胞长节毛，上部及花序分枝上的毛稠密。基生叶及下部茎叶不裂，椭圆形或长椭圆形，长6—11厘米，宽3—5厘米，基部渐狭，有翼柄或无柄；中上部茎叶与基生叶及下部茎叶同形，但较小或披针形，无柄或有短翼柄；接花序下部的常线形；全部叶顶端急尖或渐尖，边缘有稀疏的凹齿，齿顶及齿缘有小尖头，但上部的叶通常全缘，无锯齿，两面被稠密或稀疏的黑色多细胞长节毛。头状花序下垂或歪斜，通常2一?枚在茎枝顶端排成伞房花序或聚伞花序。总苞宽钟状，长1．5厘米，宽1．5—2．5厘米；总苞片3层，外层长椭圆形，长8毫米，宽1毫米，中层披针形，长ll毫米，宽1．5毫米，内层披针形或长椭圆形，长15毫米，宽2毫米；全部总苞片外面沿中脉被黑色或黑褐色多细胞长节毛，但向内层的毛较稀疏，外层顶端渐尖，中内层顶端急尖。舌状小花黄色。瘦果近纺锤形，压扁，上部黄色，下部黑色，长约7毫米，有多数(12—17条)，不等粗纵肋，顶端截形。冠毛淡黄色，2层，易折断，长2厘米，微细锯齿。花果期8—10月。
产地分布：分布云南西北部(丽江、贡山、德钦)、西藏(林芝、错那、察隅、墨脱)、四川西南部(木里、盐源)。生于高山林缘、林下、草甸或灌丛中，海拔3 200—4 200米。尼泊尔、 锡金、不丹、印度北部及缅甸北部有分布。模式标本采自尼泊尔。
参考文献：Dubyaea hispida (D．Don) DC．，Prodr．7：247．1838．Quoad nomen．；Stebbins in Mem．Torrey Bot．Club．19：18．1940．P．P．；S．Y．Hu in Quart．Journ．Taiwan Mus．19(3—4)：236．1966．P．P．；Kitam．Et 5．Gould．，Emumer．Flowering Plant Nepal 3：25，1982；西藏植物志4：954．1985．Quoadnomen．；Shih in Act．Phytotax． Sin．31：436．1993．——Hieracium hispidum D．Don，Prodr．F1．Nepal 165．1825．
《Flora of China》 Vol. 20-21 (2011)
Dubyaea hispida Candolle Prodr. 7: 247. 1838.
厚喙菊 hou hui ju
Hieracium hispidum D. Don, Prodr. Fl. Nepal. 165. 1825, not Forssk?l (1775); Crepis bhotanica Hutchinson; C. dubyaea (C. B. Clarke) C. Marquand & Airy Shaw; Dubyaea bhotanica (Hutchinson) C. Shih; D. lanceolata C. Shih; D. pteropoda C. Shih; Lactuca dubyaea C. B. Clarke.
Herbs 20-60 cm tall, perennial, with creeping ?rhizomes. Stem erect, branched from below middle or higher up, leafy, with long blackish stiff glandular hairs and pale curly weak hairs. Basal and lower stem leaves sessile or base attenuate into a long petiole-like portion, oblanceolate, elliptic, or narrowly elliptic, 5-18 × 1.5-6 cm, sinuate-dentate to shallowly lyrately pinnatifid, both faces with hairs or mainly abaxially, base narrow to widened and ± clasping, margin mucronulately dentate, apex acute to acuminate. Middle and upper stem leaves similar to lower stem leaves but ± lanceolate, smaller, base auriculately clasping; uppermost stem leaves linear-lanceolate, reduced in size. Synflorescence corymbiform, with 2-7 capitula. Capitula nodding, with 40-50 florets; peduncle blackish pilose. Involucre campanulate, 1.8-2 × 1-1.5 cm. Phyllaries with long dark brown to blackish stiff glandular hairs on midvein except on innermost ones; outer phyllaries rather few, narrowly lanceolate to linear, closely approaching inner ones in length, apex acute to acuminate; inner phyllaries many, lanceolate, margin apically often ± brownish ciliate to fringed. Florets yellow. Achene brown, 7-9 mm, with several ribs, apex long attenuate and paler. Pappus yellowish, 0.8-1.2 cm. Fl. and fr. Jul-Nov. 2n = 16.
Forests, forest margins, meadows, thickets; 2700-4500 m. Sichuan, Xizang, Yunnan [Bhutan, India (Sikkim), N Myanmar, Nepal].
Dubyaea hispida is here considered in the wider sense established by Stebbins (Mem. Torrey Bot. Club 19(3): 19. 1940) and taken up by X. Zhuang (Fl. Yunnan. 13: 704. 2004), who consequently included also the segregates D. lanceolata and D. pteropoda, distinguished only by minor differences in leaf shape. The underground parts are usually described as creeping rhizomes from which the stems arise; no specimen, however, has been seen where a true rhizome is preserved. The underground parts preserved in the material seen could also be interpreted as root-borne shoots.Dubyaea hispida is here considered in the wider sense established by Stebbins (Mem. Torrey Bot. Club 19(3): 19. 1940) and taken up by X. Zhuang (Fl. Yunnan. 13: 704. 2004), who consequently included also the segregates D. lanceolata and D. pteropoda, distinguished only by minor differences in leaf shape. The underground parts are usually described as creeping rhizomes from which the stems arise; no specimen, however, has been seen where a true rhizome is preserved. The underground parts preserved in the material seen could also be interpreted as root-borne shoots.