大叶假冷蕨Pseudocystopteris atkinsonii (Bedd.) Ching
大叶假冷蕨Pseudocystopteris atkinsonii (Bedd.) Ching
1. 大叶假冷蕨（植物分类学报）假冷蕨（台湾植物志）、阿金蹄盖蕨（中国主要植物图说 蕨类植物门）、喜马拉雅假冷蕨（植物分类学报）、大叶蹄盖蕨（福建植物志）、亚德氏蹄盖蕨（台湾植物志）图版17：1-2
Pseudocystopteris atkinsonii (Bedd.) Ching in Acta Phytotax. Sin. 9 (1): 78. 1964; Ic. Corm. Sin. 1: 176, f. 352. 1972; Fl. Tsinling. 2: 92. 1974; Y. L. Chang et al., Sporae Pterid, Sin. 237, t. 40, f. 32-33. 1976; DeVol et C. M. Kuo in H. L. Li et al. , Fl. Taiwan 1: 147, pl. 167. 1975; 河南植物志1: 51. 1981; Ching et S. K. Wu in C. Y. Wu, Fl. Xizang. 1: 126. 1983; B. Bartholomew et al. in Journ. Arnold Arbor. 64: 24. 1983; Jarrett, Ind. Fil. Suppl. 5: 163. 1985; Z. R. Wang in W. T. Wang, Vase. Pl. Hengduan Mount. 1: 76. 1993; Shing in J. F. Cheng et G. F. Zhu, Fl. Jiangxi 1: 152-153, f. 136. 1993; S. F. Wu in Journ. Wuhan Bot. Res. 12 (4): 330. 1994 et in W. T. Wang, Keys Vasc. Pl. Wuling Mount. 34. 1995. ——Athyrium atkinsonii Bedd. , Ferns Brit. S. Ind. Suppl. II, t. 359. 1876 et Handb. Ferns Brit. Ind. 163. 1883; C. Chr. , Ind. Fil. 140. 1906 et Suppl. 3: 40. 1934; H. Ito, Fil. Jap. Ill. t. 197. 1944; 傅书遐, 中国主要植物图说 蕨类植物门113, 图146. 1957; Ohwi, Fl. Jap. Pterid. 112. 1957; Tagawa, Col. Ill. Jap. Pterid. 128, 180, t. 51, f. 281. 1959; Namegata et Kurata, Enum. Jap. Pterid. 276. 1961; Nakaike, Enum. Pterid. Jap. 129. 1975 et New Fl. Jap. Pterid. 274, cum f. 1982 et 1. c. Rev. Enl. 1992; Fl. Fujian. 1: 114. 1982; Y. J. Ling et al. in T. W, Liu, Fl. Shanxi. 1: 69, f. 35. 1992; W. C. Shieh et al. in T. C. Huang, Fl. Taiwan 2nd ed. 1: 418, Pl. 168. 1994. ——Asplenium atkinsonii Clarke in Trans. Linn. Soc. , Lond. , ser. 2, Bot. 1: 487, t. 57. 1880. ——Polypodium foliolosum Wall. , nom. nud. ——Asplenium andersonii Clarke in Journ. Linn. Soc. 15: 154. 1876. ——Asplenium atkinsonii Bedd. var. andersonii Clarke in Trans. Linn. Soc. 2. Bot. 1: 487, t. 52. 1880. ——Pseudocystopteris andersonii (Clarke) Ching in Acta Phytotax. Sin. 9: 79. 1964. ——Asplenium fimbriatum Wall, var. sphaeropteroides Clarke in Trans. Linn. Soc. , Lond. , 2 Bot. 1: 495, excl. t. 62, f. 1. 1880. ——Athyrium fimbriatum var. sphaeropteroides Bedd. , Handb. Ferns Brit. India. 172. 1883. ——Aspidium senanense Franch. et Sav. , Enum. Pl. Jap. 2: 241. 1876 et 636. 1879. ——Dryopteris senanensis C. Chr. , Ind. Fil. 291. 1906; Makino et Nemoto, Fl. Jap. 1623. 1925. ——Athyrium senanense Koidz. et Tagawa in Tagawa in Acta Phytotax. Geobot. 3: 34. 1934. ——Asplenium lastreoides Bak. in Journ. Bot. 227. 1888. ——Athyrium lastreoides Diels in Engl. u. Prantl Nat. Pflanzenfam. 1 (4): 224. 1899; C. Chr. , Ind. Fil. 143. 1906. ——Davallia athyrifolia Bak. in Ann. Bot. 5: 200. 1891——Polypodium leptophyllum Bak. in Ann. Bot. 5: 458. 1891, non L. 1753, nec Moritz ex Mett. 1864. ——Dryopteris gracilifrons C. Chr. , Ind. Fil. 268. 1906, nom. nov. for Polypodium leptophyllum Bak. ——Leucostegia yaklaensis Bedd. , Suppl. Ferns Brit. Ind. Ceylon. 13-14. 1892. ——Davallia yaklaensis C. Chr. , Ind. Fil. 215. 1906. ——Araiostegia yaklaensis Nayar et Kaur, Comp. Bedd. Handb. 17. 1974. ——Athyrium microsorum Makino in Bot. Mag. Tokyo 13: 25. 1899; Matsum. Ind. Pl. Jap. 294. 1904. ——Athyrium filix-femina Roth var. deltoidea Christ in Bull. Soc. Bot. France 52. Mem. 1: 46. 1905. ——Aspidium fargesii Christ in Bull. Soc. France 52 Mem. 1: 42. 1905. ——Dryopteris fargesii C. Chr. , Ind. Fil. 264. 1906. ——Athyrium monticola Rosenst. in Hedwigia 13: 113. 1913; C. Chr. , Ind. Fil. Suppl. 2: 7. 1917. ——Cystopteris grandis C. Chr. in Lev. Cat. Pl. Yun-Nan 100. 1916.
根状茎粗而横卧，直径0.5-1厘米，带有残留的叶柄基部，先端及嫩叶叶柄基部被有较多的浅褐色或深褐色卵状鳞片；叶近生或远生。能育叶长 (15-) 60-100 (-200)厘米；叶柄长20-50 (-100)厘米，基部黑褐色，直径3-10毫米，向上渐变为禾秆色或稍带淡紫红色，偶有细鳞片；叶片通常较大，长(24-) 35-50 (-100)厘米，宽与长几相等或稍狭，阔卵形或卵状三角形，小植株的叶往往卵形，而大植株的叶则近三角形，先端渐尖，成熟叶片二回羽状至四回羽状，叶轴和羽轴稍曲折；羽片10-12 (-20)对，互生，斜向上，X0.5-1 (-3) 厘米长的短柄，彼此离开、接近或略呈覆瓦状，基部一对羽片和第二对同形，相距5-15厘米，长15-20 (-50)厘米，中部宽可达6-8 (-28)厘米，基部略变狭，长卵形至长卵状披针形，渐尖头，基部近平截，二回羽裂至三回羽状；一回小羽片10对或更多，上先出，互生，有长约2毫米的短柄，相距1.4-2厘米，近平展，基部一对略缩短，中部的长3.5-5厘米，宽1.2-2厘米，长圆状披针形，渐尖头，基部平截，一回羽状二回小羽片羽裂至二回羽状；二回小羽片6-8对，长5-8毫米，宽2-3毫米，长圆形或近卵形，先端钝圆，并有不整齐的短锯齿，基部阔楔形，下侧略下延，一回羽状或羽裂；末回裂片卵状长圆形，先端圆钝或近方头，并有2-3个不整齐的短锯齿，基部阔楔形，略不对称。叶脉两面可见，下面明显，在末回小羽片上有羽状侧脉4-5对，单一，偶有二叉。叶干后草质，黑褐色或褐绿色，两面无毛，主脉或小羽轴上面有间断的槽状隆起，沿叶轴及各回小羽轴下面偶有膜质、披针形小鳞片。孢子囊群圆形或椭圆形，背生或半侧生于裂片基部上侧之小脉上，每裂片通常一枚，每末回小羽片通常2-3对；囊群盖小，通常圆肾形，少有长圆形或弯钩形，膜质，灰褐色，边缘略啮蚀，易脱落，成熟的孢子囊群往往好象无盖。孢子周壁表面具褶皱，凹凸不平，形成拟网状。染色体数目n=40。
分布于山西南部（垣曲）、陕西（太白山、宁陕、平利、佛坪）、甘肃（康县、徽县）、江西（武功山、安福、井冈山）、福建（崇安）、台湾（南投、嘉义）、河南（伏牛山、大别山、老君山、卢氏、桐柏、嵩县、奕川）、湖北（神农架）、湖南（桑植、石门）、四川（峨眉山、都江堰、保安、石绵、甘孜、马尔康、道孚、木里、盐源、康定、雷波、九龙、越西、大相岭、金川、雅江）、重庆（城口、开县）、贵州（梵净山、江口、威宁）、云南（鹤庆、丽江、泸水、贡山、漾濞、中甸、维西、德钦、宾川、昆明、禄劝、呈贡、嵩明、宜良、会泽、澄江、大姚、巧家、大关、昭通、宁浪、大理）和西藏东南部（察隅、吉隆、墨脱、错那）等省区，生冷杉或铁杉林下或灌丛中阴湿处，海拔1200-4000米。日本、韩国、缅甸北部、锡金、不丹、尼泊尔、印度、巴基斯坦北部和克什米尔地区均有分布。模式标本采自锡金。
《Flora of China》 Vol. 2-3 (2013)
Athyrium atkinsonii Beddome Suppl. Ferns S. Ind. 11. 1876.
大叶假冷蕨 da ye jia leng jue
Araiostegia yaklaensis (Beddome) B. K. Nayar & S. Kaur; Aspidium senanense Franchet & Savatier; Asplenium andersonii C. B. Clarke; A. atkinsonii (Beddome) C. B. Clarke; A. atkinsonii var. andersonii (C. B. Clarke) C. B. Clarke; A. fimbriatum Kunze var. sphaeropteroides C. B. Clarke; A. lastreoides Baker; Athyrium andersonii (C. B. Clarke) Panigrahi & Basu; A. filix-femina (Linnaeus) Roth var. deltoidea Christ; A. fimbriatum Hooker ex T. Moore var. sphaeropteroides (C. B. Clarke) Beddome; A. lastreoides (Baker) Diels; A. microsorum Makino; A. monticola Rosenstock; A. senanense (Franchet & Savatier) Koidzumi & Tagawa; Cystopteris grandis C. Christensen; Davallia athyriifolia Baker; D. yaklaensis (Beddome) C. Christensen; Dryopteris gracilifrons C. Christensen; D. senanensis (Franchet & Savatier) C. Christensen; Leucostegia yaklaensis Beddome; Polypodium leptophyllum Baker (1891), not Linnaeus (1753); Pseudocystopteris andersonii (C. B. Clarke) Ching; P. atkinsonii (Beddome) Ching.
Rhizomes shortly creeping, 0.5-1 cm in diam., bearing remaining stipe bases, apex together with stipe bases, clothed with brownish or brown ovate scales. Fronds approximate or distant; fertile fronds (15-)60-100(-200) cm; stipe blackish brown at base, upper part stramineous or pale purplish red, 20-50(-100) cm, 3-10 mm in diam., occasionally with thin scales; lamina (24-)35-50(-100) cm, nearly as broad as long or slightly narrower, broadly ovate or ovate-triangular, in small plants lamina often ovate, in large plants lamina often subtriangular, 2-4-pinnate, acuminate; rachis and costae slightly tortuous; pinnae 10-12(-20) pairs, alternate, ascending, stalk 0.5-1(-3) cm, separate from each other, sometimes imbricate; basal pinnae as large as second basal pinnae, 5-15 cm apart, 15-20(-50) × up to 6-8(-28) cm at middle, slightly narrowed toward base, narrowly ovate to narrowly ovate-lanceolate, acuminate, subtruncate at base, 2-pinnate-pinnatifid to 3-pinnate; pinnules 10 pairs or more, anadromous, alternate, with stalk ca. 2 mm, 1.4-2 cm apart, subspreading, basal pair of pinnules slightly shortened, middle pinnules oblong-lanceolate, 3.5-5 × 1.2-2 cm, acuminate, truncate at base; secondary pinnules 6-8 pairs, oblong or subovate, 5-8 × 2-3 mm, obtuse-rounded, crenate at apex, base broadly cuneate, basiscopic base slightly decurrent; ultimate segments ovate-oblong, rounded-obtuse, or subsquare, with 2 or 3 unequal short teeth at apex, broadly cuneate and slightly inequilateral at base. Veins visible on surfaces, more prominent abaxially, pinnate with 4 or 5 pairs of lateral veins, simple, rarely forked. Lamina herbaceous when dried, blackish brown or greenish brown, glabrate; adaxial groove of costule open to groove of midribs, rachis and costae occasionally with membranous, small-lanceolate scales abaxially, without spines at base of pinnules and segments. Sori orbicular or elliptic, abaxial or semilateral on basal acroscopic veins; indusia pale brown, small, orbicular-reniform, occasionally oblong, or J-shaped, membranous, slightly erose at margin, fugacious, apparently exindusiate when mature. Perispore folded. 2n = 80.
Damp areas in coniferous forests and shrubs; 1200-4000 m. Chongqing, Fujian, Gansu, Guizhou, Henan, Hubei, Hunan, Jiangxi, S Shaanxi, Shanxi, Sichuan, Taiwan, SE Xizang, Yunnan [Bhutan, Himalaya, N India, Japan, Kashmir, S Korea, N Myanmar, Nepal, N Pakistan].
“Polypodium foliolosum” (Wallich, Numer. List, no. 318. 1829) belongs here but is a nomen nudum and was not therefore validly published (Melbourne Code, Art. 38.1(a)).
别名：假冷蕨;阿金蹄盖蕨;喜马拉雅假冷蕨;大叶蹄盖蕨;大假冷蕨;亚德氏蹄盖蕨;
科名：蹄盖蕨科 Athyriaceae
属名：假冷蕨属 Pseudocystopteris
1. 大叶假冷蕨（植物分类学报）假冷蕨（台湾植物志）、阿金蹄盖蕨（中国主要植物图说 蕨类植物门）、喜马拉雅假冷蕨（植物分类学报）、大叶蹄盖蕨（福建植物志）、亚德氏蹄盖蕨（台湾植物志）图版17：1-2
Pseudocystopteris atkinsonii (Bedd.) Ching in Acta Phytotax. Sin. 9 (1): 78. 1964; Ic. Corm. Sin. 1: 176, f. 352. 1972; Fl. Tsinling. 2: 92. 1974; Y. L. Chang et al., Sporae Pterid, Sin. 237, t. 40, f. 32-33. 1976; DeVol et C. M. Kuo in H. L. Li et al. , Fl. Taiwan 1: 147, pl. 167. 1975; 河南植物志1: 51. 1981; Ching et S. K. Wu in C. Y. Wu, Fl. Xizang. 1: 126. 1983; B. Bartholomew et al. in Journ. Arnold Arbor. 64: 24. 1983; Jarrett, Ind. Fil. Suppl. 5: 163. 1985; Z. R. Wang in W. T. Wang, Vase. Pl. Hengduan Mount. 1: 76. 1993; Shing in J. F. Cheng et G. F. Zhu, Fl. Jiangxi 1: 152-153, f. 136. 1993; S. F. Wu in Journ. Wuhan Bot. Res. 12 (4): 330. 1994 et in W. T. Wang, Keys Vasc. Pl. Wuling Mount. 34. 1995. ——Athyrium atkinsonii Bedd. , Ferns Brit. S. Ind. Suppl. II, t. 359. 1876 et Handb. Ferns Brit. Ind. 163. 1883; C. Chr. , Ind. Fil. 140. 1906 et Suppl. 3: 40. 1934; H. Ito, Fil. Jap. Ill. t. 197. 1944; 傅书遐, 中国主要植物图说 蕨类植物门113, 图146. 1957; Ohwi, Fl. Jap. Pterid. 112. 1957; Tagawa, Col. Ill. Jap. Pterid. 128, 180, t. 51, f. 281. 1959; Namegata et Kurata, Enum. Jap. Pterid. 276. 1961; Nakaike, Enum. Pterid. Jap. 129. 1975 et New Fl. Jap. Pterid. 274, cum f. 1982 et 1. c. Rev. Enl. 1992; Fl. Fujian. 1: 114. 1982; Y. J. Ling et al. in T. W, Liu, Fl. Shanxi. 1: 69, f. 35. 1992; W. C. Shieh et al. in T. C. Huang, Fl. Taiwan 2nd ed. 1: 418, Pl. 168. 1994. ——Asplenium atkinsonii Clarke in Trans. Linn. Soc. , Lond. , ser. 2, Bot. 1: 487, t. 57. 1880. ——Polypodium foliolosum Wall. , nom. nud. ——Asplenium andersonii Clarke in Journ. Linn. Soc. 15: 154. 1876. ——Asplenium atkinsonii Bedd. var. andersonii Clarke in Trans. Linn. Soc. 2. Bot. 1: 487, t. 52. 1880. ——Pseudocystopteris andersonii (Clarke) Ching in Acta Phytotax. Sin. 9: 79. 1964. ——Asplenium fimbriatum Wall, var. sphaeropteroides Clarke in Trans. Linn. Soc. , Lond. , 2 Bot. 1: 495, excl. t. 62, f. 1. 1880. ——Athyrium fimbriatum var. sphaeropteroides Bedd. , Handb. Ferns Brit. India. 172. 1883. ——Aspidium senanense Franch. et Sav. , Enum. Pl. Jap. 2: 241. 1876 et 636. 1879. ——Dryopteris senanensis C. Chr. , Ind. Fil. 291. 1906; Makino et Nemoto, Fl. Jap. 1623. 1925. ——Athyrium senanense Koidz. et Tagawa in Tagawa in Acta Phytotax. Geobot. 3: 34. 1934. ——Asplenium lastreoides Bak. in Journ. Bot. 227. 1888. ——Athyrium lastreoides Diels in Engl. u. Prantl Nat. Pflanzenfam. 1 (4): 224. 1899; C. Chr. , Ind. Fil. 143. 1906. ——Davallia athyrifolia Bak. in Ann. Bot. 5: 200. 1891——Polypodium leptophyllum Bak. in Ann. Bot. 5: 458. 1891, non L. 1753, nec Moritz ex Mett. 1864. ——Dryopteris gracilifrons C. Chr. , Ind. Fil. 268. 1906, nom. nov. for Polypodium leptophyllum Bak. ——Leucostegia yaklaensis Bedd. , Suppl. Ferns Brit. Ind. Ceylon. 13-14. 1892. ——Davallia yaklaensis C. Chr. , Ind. Fil. 215. 1906. ——Araiostegia yaklaensis Nayar et Kaur, Comp. Bedd. Handb. 17. 1974. ——Athyrium microsorum Makino in Bot. Mag. Tokyo 13: 25. 1899; Matsum. Ind. Pl. Jap. 294. 1904. ——Athyrium filix-femina Roth var. deltoidea Christ in Bull. Soc. Bot. France 52. Mem. 1: 46. 1905. ——Aspidium fargesii Christ in Bull. Soc. France 52 Mem. 1: 42. 1905. ——Dryopteris fargesii C. Chr. , Ind. Fil. 264. 1906. ——Athyrium monticola Rosenst. in Hedwigia 13: 113. 1913; C. Chr. , Ind. Fil. Suppl. 2: 7. 1917. ——Cystopteris grandis C. Chr. in Lev. Cat. Pl. Yun-Nan 100. 1916.
根状茎粗而横卧，直径0.5-1厘米，带有残留的叶柄基部，先端及嫩叶叶柄基部被有较多的浅褐色或深褐色卵状鳞片；叶近生或远生。能育叶长 (15-) 60-100 (-200)厘米；叶柄长20-50 (-100)厘米，基部黑褐色，直径3-10毫米，向上渐变为禾秆色或稍带淡紫红色，偶有细鳞片；叶片通常较大，长(24-) 35-50 (-100)厘米，宽与长几相等或稍狭，阔卵形或卵状三角形，小植株的叶往往卵形，而大植株的叶则近三角形，先端渐尖，成熟叶片二回羽状至四回羽状，叶轴和羽轴稍曲折；羽片10-12 (-20)对，互生，斜向上，X0.5-1 (-3) 厘米长的短柄，彼此离开、接近或略呈覆瓦状，基部一对羽片和第二对同形，相距5-15厘米，长15-20 (-50)厘米，中部宽可达6-8 (-28)厘米，基部略变狭，长卵形至长卵状披针形，渐尖头，基部近平截，二回羽裂至三回羽状；一回小羽片10对或更多，上先出，互生，有长约2毫米的短柄，相距1.4-2厘米，近平展，基部一对略缩短，中部的长3.5-5厘米，宽1.2-2厘米，长圆状披针形，渐尖头，基部平截，一回羽状二回小羽片羽裂至二回羽状；二回小羽片6-8对，长5-8毫米，宽2-3毫米，长圆形或近卵形，先端钝圆，并有不整齐的短锯齿，基部阔楔形，下侧略下延，一回羽状或羽裂；末回裂片卵状长圆形，先端圆钝或近方头，并有2-3个不整齐的短锯齿，基部阔楔形，略不对称。叶脉两面可见，下面明显，在末回小羽片上有羽状侧脉4-5对，单一，偶有二叉。叶干后草质，黑褐色或褐绿色，两面无毛，主脉或小羽轴上面有间断的槽状隆起，沿叶轴及各回小羽轴下面偶有膜质、披针形小鳞片。孢子囊群圆形或椭圆形，背生或半侧生于裂片基部上侧之小脉上，每裂片通常一枚，每末回小羽片通常2-3对；囊群盖小，通常圆肾形，少有长圆形或弯钩形，膜质，灰褐色，边缘略啮蚀，易脱落，成熟的孢子囊群往往好象无盖。孢子周壁表面具褶皱，凹凸不平，形成拟网状。染色体数目n=40。
分布于山西南部（垣曲）、陕西（太白山、宁陕、平利、佛坪）、甘肃（康县、徽县）、江西（武功山、安福、井冈山）、福建（崇安）、台湾（南投、嘉义）、河南（伏牛山、大别山、老君山、卢氏、桐柏、嵩县、奕川）、湖北（神农架）、湖南（桑植、石门）、四川（峨眉山、都江堰、保安、石绵、甘孜、马尔康、道孚、木里、盐源、康定、雷波、九龙、越西、大相岭、金川、雅江）、重庆（城口、开县）、贵州（梵净山、江口、威宁）、云南（鹤庆、丽江、泸水、贡山、漾濞、中甸、维西、德钦、宾川、昆明、禄劝、呈贡、嵩明、宜良、会泽、澄江、大姚、巧家、大关、昭通、宁浪、大理）和西藏东南部（察隅、吉隆、墨脱、错那）等省区，生冷杉或铁杉林下或灌丛中阴湿处，海拔1200-4000米。日本、韩国、缅甸北部、锡金、不丹、尼泊尔、印度、巴基斯坦北部和克什米尔地区均有分布。模式标本采自锡金。
《Flora of China》 Vol. 2-3 (2013)
Athyrium atkinsonii Beddome Suppl. Ferns S. Ind. 11. 1876.
大叶假冷蕨 da ye jia leng jue
Araiostegia yaklaensis (Beddome) B. K. Nayar & S. Kaur; Aspidium senanense Franchet & Savatier; Asplenium andersonii C. B. Clarke; A. atkinsonii (Beddome) C. B. Clarke; A. atkinsonii var. andersonii (C. B. Clarke) C. B. Clarke; A. fimbriatum Kunze var. sphaeropteroides C. B. Clarke; A. lastreoides Baker; Athyrium andersonii (C. B. Clarke) Panigrahi & Basu; A. filix-femina (Linnaeus) Roth var. deltoidea Christ; A. fimbriatum Hooker ex T. Moore var. sphaeropteroides (C. B. Clarke) Beddome; A. lastreoides (Baker) Diels; A. microsorum Makino; A. monticola Rosenstock; A. senanense (Franchet & Savatier) Koidzumi & Tagawa; Cystopteris grandis C. Christensen; Davallia athyriifolia Baker; D. yaklaensis (Beddome) C. Christensen; Dryopteris gracilifrons C. Christensen; D. senanensis (Franchet & Savatier) C. Christensen; Leucostegia yaklaensis Beddome; Polypodium leptophyllum Baker (1891), not Linnaeus (1753); Pseudocystopteris andersonii (C. B. Clarke) Ching; P. atkinsonii (Beddome) Ching.
Rhizomes shortly creeping, 0.5-1 cm in diam., bearing remaining stipe bases, apex together with stipe bases, clothed with brownish or brown ovate scales. Fronds approximate or distant; fertile fronds (15-)60-100(-200) cm; stipe blackish brown at base, upper part stramineous or pale purplish red, 20-50(-100) cm, 3-10 mm in diam., occasionally with thin scales; lamina (24-)35-50(-100) cm, nearly as broad as long or slightly narrower, broadly ovate or ovate-triangular, in small plants lamina often ovate, in large plants lamina often subtriangular, 2-4-pinnate, acuminate; rachis and costae slightly tortuous; pinnae 10-12(-20) pairs, alternate, ascending, stalk 0.5-1(-3) cm, separate from each other, sometimes imbricate; basal pinnae as large as second basal pinnae, 5-15 cm apart, 15-20(-50) × up to 6-8(-28) cm at middle, slightly narrowed toward base, narrowly ovate to narrowly ovate-lanceolate, acuminate, subtruncate at base, 2-pinnate-pinnatifid to 3-pinnate; pinnules 10 pairs or more, anadromous, alternate, with stalk ca. 2 mm, 1.4-2 cm apart, subspreading, basal pair of pinnules slightly shortened, middle pinnules oblong-lanceolate, 3.5-5 × 1.2-2 cm, acuminate, truncate at base; secondary pinnules 6-8 pairs, oblong or subovate, 5-8 × 2-3 mm, obtuse-rounded, crenate at apex, base broadly cuneate, basiscopic base slightly decurrent; ultimate segments ovate-oblong, rounded-obtuse, or subsquare, with 2 or 3 unequal short teeth at apex, broadly cuneate and slightly inequilateral at base. Veins visible on surfaces, more prominent abaxially, pinnate with 4 or 5 pairs of lateral veins, simple, rarely forked. Lamina herbaceous when dried, blackish brown or greenish brown, glabrate; adaxial groove of costule open to groove of midribs, rachis and costae occasionally with membranous, small-lanceolate scales abaxially, without spines at base of pinnules and segments. Sori orbicular or elliptic, abaxial or semilateral on basal acroscopic veins; indusia pale brown, small, orbicular-reniform, occasionally oblong, or J-shaped, membranous, slightly erose at margin, fugacious, apparently exindusiate when mature. Perispore folded. 2n = 80.
Damp areas in coniferous forests and shrubs; 1200-4000 m. Chongqing, Fujian, Gansu, Guizhou, Henan, Hubei, Hunan, Jiangxi, S Shaanxi, Shanxi, Sichuan, Taiwan, SE Xizang, Yunnan [Bhutan, Himalaya, N India, Japan, Kashmir, S Korea, N Myanmar, Nepal, N Pakistan].
“Polypodium foliolosum” (Wallich, Numer. List, no. 318. 1829) belongs here but is a nomen nudum and was not therefore validly published (Melbourne Code, Art. 38.1(a)).