光蹄盖蕨Athyrium otophorum (Miq.) Koidz.
光蹄盖蕨Athyrium otophorum (Miq.) Koidz.
93. 光蹄盖蕨（中国蕨类植物图谱）缙云山蹄盖蕨（中国蕨类植物孢子形态）、禾秆色蹄盖蕨（植物研究）、红足蹄盖蕨（西北植物学报）图版55：1-4
Athyrium otophorum (Miq.) Koidz., Fl. Symb. Orient. -Asiat. 40. 1930; C. Chr., Ind. Fil. Suppl. 3: 43. 1934; Ching, Ic. Fil. Sin. 3: t. 109. 1935; H. Ito, Fil. Jap. Ill. t. 180. 1944; 傅书遐, 中国主要植物图说 蕨类植物门110, 图140. 1957; Ohwi, Fl. Jap. Pterid. 121. 1957 et Fl. Jap. 85. 1965; Tagawa, Col. Ill. Jap. Pterid. 125, 185, t. 49, f. 274. 1959; Ic. Corm. Sin. 1: 181, f. 361. 1972; Nakaike, Enum. Pterid. Jap. 140. 1975 et New Fl. Jap. Pterid. 296, t. 296. 1982 et l. c. Rev. Enl. 1992; Fl. Fujian. 1: 113, t. 102. 1982; C. M. Kuo in Taiwania 30: 34, 65. 1985; C. F. Zhang in C. F. Zhang et S. Y. Zhang, Fl. Zhejiang 1: 141, f. 1-147. 1993; S. F. Wu in W. T. Wang, Keys Vase. Pl. Wuling Mount. 31. 1995. ——Asplenium otophorum Miq. in Ann. Mus. Bot. Lugd. Bat. 3: 175. 1867; Franch. et Sav., Enum. Pl. Jap. 2: 229. 1876. ——Diplazium otophorum C. Chr., Ind. Fil. 236. 1906. ——Athyrium rigescens Makino in Bot. Mag. Tokyo 13: 27. 1899; Ogata, Ic. Fil. Jap. 5: t. 216. 1933. ——Athyrium wardii Christ in Bull. Soc. Bot. France 52 Mem. 1: 49. 1905 non Makino 1857. ——Athyrium nigripes sensu Christ, Bull. Acad. Geogr. Bot. Mans 174. 1909. ——Athyrium violascens Diels in Engl. Bot. Jahrb. 29: 196. 1900; C. Chr., Ind. Fil. 147. 1906 et in Acta Hort. Gothob. 1: 76. 1924. ——Athyrium vidalii Nakai var. chinense Koidz., Fl. Symb. Orient. -Asiat. 39. 1930. ——Athyrium tsingyingense Ching in Y. L. Chang et al., Sporae Pterid. Sin. 208, t. 41, f. 17-18. 1976, nom. nud. -——Athyrium sinovidalii (sino-vidalii) Ching et Z. Y. Liu in Bull. Bot. Res. 4 (3): 8-9, t. 10. 1984. ——Athyrium erythrocaulon Ching in Ching et Hsieh in Acta Bot. Bor. -Occ. Sin. 6 (3): 154. 1986.
根状茎短，先端斜升，密被深褐色或黑褐色、线状披针形、先端纤维状的鳞片；叶簇生。能育叶长45-70(-85)厘米；叶柄长(15-)25-35厘米，直径2.5-3毫米，基部黑褐色，密被与根状茎上同样的鳞片，向上略带淡紫红色，光滑；叶片长卵形或三角状卵形，长25-35(-50)厘米，中部宽20-25厘米，先端急狭缩，基部不变狭，二回羽状；羽片约15对，急狭缩部以下约有7对，基部的对生，向上的互生，几平展，无柄或有极短柄，披针形，中部的长10-12(-25)厘米，宽2.5-3.5厘米，先端长渐尖，基部截形，一回羽状；小羽片14-17对，互生，几平展，无柄，下部的近三角形至长圆状披针形，长1-1.7厘米，中部宽4-6毫米，尖头，基部不对称，上侧截形，并有三角形的耳状凸起，与羽轴并行，下侧楔形，近全缘或上侧边缘有小锯齿。叶脉下面明显，上面不见，在小羽片上为羽状，侧脉7-8对，斜向上，下部的分叉，基部上侧的2脉为羽状或三叉。叶干后纸质，浅褐色，两面无毛；叶轴和羽轴下面淡紫红色，光滑，上面沿沟两侧边上有贴伏的钻状短硬刺。孢子囊群长圆形或短线形，每小羽片3-5对，生于叶边与主脉中间，稍近主脉；囊群盖同形，浅褐色，膜质，全缘，宿存。孢子周壁表面无褶皱，有颗粒状纹饰。然色体数目n=c. 80，2n=160。
分布于安徽、浙江（淳安、临安）、福建（崇安、建阳、将乐、龙栖山）、台湾、湖北（鹤峰、宣恩）、湖南（凤凰、桑植、永顺、沅陵、张家界）、广东（乳源）、广西（花坪、环江、九万山、隆林金钟山、龙胜、天峨、阳山、瑶山）、四川（峨眉山）、重庆（缙云山、南川金佛山）、贵州（贵定云雾、贵阳、凯里、荔波、三合、松桃、望谟、印江、江口梵净山）和云南（广南）。生常绿阔叶林或竹林下阴湿处，海拔400-1400(-2100)米。日本和朝鲜半岛也有分布。模式标本采自日本。
《Flora of China》 Vol. 2-3 (2013)
Athyrium otophorum (Miquel) Koidzumi Fl. Symb. Orient.-Asiat. 40. 1930.
光蹄盖蕨 guang ti gai jue
Asplenium otophorum Miquel, Ann. Mus. Bot. Lugduno-Batavi 3: 175. 1867; Athyrium erythrocaulon Ching; A. rigescens Makino; A. sinovidalii Ching & Z. Y. Liu; A. vidalii Nakai var. chinense Koidzumi; A. violascens Diels; Diplazium otophorum (Miquel) C. Christensen.
Rhizomes short, ascending, apex densely clothed with brown or blackish brown, linear-lanceolate, fibriform-tipped scales. Fronds caespitose; fertile fronds 45-70(-85) cm; stipe blackish brown, upward slightly purplish red, (15-)25-35 cm, 2.5-3 mm in diam., base densely clothed with scales, upward glabrate; lamina 2-pinnate, narrowly ovate or deltoid-ovate, 25-35(-50) × 20-25 cm at middle, base not narrowed, apex abruptly narrowed; pinnae ca. 15 pairs, under abruptly narrowed tip ca. 7 pairs, basal pinnae opposite, upper pinnae alternate, subspreading, sessile or very shortly stalked, lanceolate; middle pinnae 10-12(-25) × 2.5-3.5 cm, base truncate, pinnate, apex long acuminate; pinnules 14-17 pairs, alternate, subspreading, sessile, lower pinnules subtriangular to oblong-lanceolate, 1-1.7 cm × 4-6 mm at middle, base inequilateral, truncate, parallel to costae, auriculate on acroscopic side, auricles triangular, cuneate on basiscopic side, subentire or with small tooth at acroscopic margin, apex acute; veins visible abaxially, inconspicuous adaxially, pinnate in pinnules, lateral veins 7 or 8 pairs, oblique, lower veins forked, basal acroscopic veins 2-pinnate or 3-forked. Lamina papery when dried, pale brown, glabrate on both surfaces; rachis and costae pale red-purple, glabrate abaxially, with subulate spines on adaxial side at base of costa or costule. Sori oblong or shortly linear, 3-5 pairs per pinnule, inframedial; indusia brownish, oblong or shortly linear, membranous, entire, persistent. Perispore surface without folds, with granular ornamentation. 2n = 160.
Evergreen broad-leaved forests, bamboo forests, wet areas; 400-1400(-3000) m. Anhui, Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Sichuan, Taiwan, Yunnan, Zhejiang [Japan, Korea].
Knapp (Ferns Fern Allies Taiwan, 439. 2011) noted that Athyrium otophorum is closely allied with A. delavayi, A. epirachis, and A. pubicostatum; and, although it is often distinguished by the glabrous abaxial surface of the rachis and costae, sessile pinnae, and auriculate pinnule bases, its diagnostic characters are not always definite.
别名：缙云山蹄盖蕨;红足蹄盖蕨;禾秆色蹄盖蕨;光秆蹄盖蕨;光柄蹄盖蕨;光蹄蹄盖蕨;
科名：蹄盖蕨科 Athyriaceae
属名：蹄盖蕨属 Athyrium
93. 光蹄盖蕨（中国蕨类植物图谱）缙云山蹄盖蕨（中国蕨类植物孢子形态）、禾秆色蹄盖蕨（植物研究）、红足蹄盖蕨（西北植物学报）图版55：1-4
Athyrium otophorum (Miq.) Koidz., Fl. Symb. Orient. -Asiat. 40. 1930; C. Chr., Ind. Fil. Suppl. 3: 43. 1934; Ching, Ic. Fil. Sin. 3: t. 109. 1935; H. Ito, Fil. Jap. Ill. t. 180. 1944; 傅书遐, 中国主要植物图说 蕨类植物门110, 图140. 1957; Ohwi, Fl. Jap. Pterid. 121. 1957 et Fl. Jap. 85. 1965; Tagawa, Col. Ill. Jap. Pterid. 125, 185, t. 49, f. 274. 1959; Ic. Corm. Sin. 1: 181, f. 361. 1972; Nakaike, Enum. Pterid. Jap. 140. 1975 et New Fl. Jap. Pterid. 296, t. 296. 1982 et l. c. Rev. Enl. 1992; Fl. Fujian. 1: 113, t. 102. 1982; C. M. Kuo in Taiwania 30: 34, 65. 1985; C. F. Zhang in C. F. Zhang et S. Y. Zhang, Fl. Zhejiang 1: 141, f. 1-147. 1993; S. F. Wu in W. T. Wang, Keys Vase. Pl. Wuling Mount. 31. 1995. ——Asplenium otophorum Miq. in Ann. Mus. Bot. Lugd. Bat. 3: 175. 1867; Franch. et Sav., Enum. Pl. Jap. 2: 229. 1876. ——Diplazium otophorum C. Chr., Ind. Fil. 236. 1906. ——Athyrium rigescens Makino in Bot. Mag. Tokyo 13: 27. 1899; Ogata, Ic. Fil. Jap. 5: t. 216. 1933. ——Athyrium wardii Christ in Bull. Soc. Bot. France 52 Mem. 1: 49. 1905 non Makino 1857. ——Athyrium nigripes sensu Christ, Bull. Acad. Geogr. Bot. Mans 174. 1909. ——Athyrium violascens Diels in Engl. Bot. Jahrb. 29: 196. 1900; C. Chr., Ind. Fil. 147. 1906 et in Acta Hort. Gothob. 1: 76. 1924. ——Athyrium vidalii Nakai var. chinense Koidz., Fl. Symb. Orient. -Asiat. 39. 1930. ——Athyrium tsingyingense Ching in Y. L. Chang et al., Sporae Pterid. Sin. 208, t. 41, f. 17-18. 1976, nom. nud. -——Athyrium sinovidalii (sino-vidalii) Ching et Z. Y. Liu in Bull. Bot. Res. 4 (3): 8-9, t. 10. 1984. ——Athyrium erythrocaulon Ching in Ching et Hsieh in Acta Bot. Bor. -Occ. Sin. 6 (3): 154. 1986.
根状茎短，先端斜升，密被深褐色或黑褐色、线状披针形、先端纤维状的鳞片；叶簇生。能育叶长45-70(-85)厘米；叶柄长(15-)25-35厘米，直径2.5-3毫米，基部黑褐色，密被与根状茎上同样的鳞片，向上略带淡紫红色，光滑；叶片长卵形或三角状卵形，长25-35(-50)厘米，中部宽20-25厘米，先端急狭缩，基部不变狭，二回羽状；羽片约15对，急狭缩部以下约有7对，基部的对生，向上的互生，几平展，无柄或有极短柄，披针形，中部的长10-12(-25)厘米，宽2.5-3.5厘米，先端长渐尖，基部截形，一回羽状；小羽片14-17对，互生，几平展，无柄，下部的近三角形至长圆状披针形，长1-1.7厘米，中部宽4-6毫米，尖头，基部不对称，上侧截形，并有三角形的耳状凸起，与羽轴并行，下侧楔形，近全缘或上侧边缘有小锯齿。叶脉下面明显，上面不见，在小羽片上为羽状，侧脉7-8对，斜向上，下部的分叉，基部上侧的2脉为羽状或三叉。叶干后纸质，浅褐色，两面无毛；叶轴和羽轴下面淡紫红色，光滑，上面沿沟两侧边上有贴伏的钻状短硬刺。孢子囊群长圆形或短线形，每小羽片3-5对，生于叶边与主脉中间，稍近主脉；囊群盖同形，浅褐色，膜质，全缘，宿存。孢子周壁表面无褶皱，有颗粒状纹饰。然色体数目n=c. 80，2n=160。
分布于安徽、浙江（淳安、临安）、福建（崇安、建阳、将乐、龙栖山）、台湾、湖北（鹤峰、宣恩）、湖南（凤凰、桑植、永顺、沅陵、张家界）、广东（乳源）、广西（花坪、环江、九万山、隆林金钟山、龙胜、天峨、阳山、瑶山）、四川（峨眉山）、重庆（缙云山、南川金佛山）、贵州（贵定云雾、贵阳、凯里、荔波、三合、松桃、望谟、印江、江口梵净山）和云南（广南）。生常绿阔叶林或竹林下阴湿处，海拔400-1400(-2100)米。日本和朝鲜半岛也有分布。模式标本采自日本。
《Flora of China》 Vol. 2-3 (2013)
Athyrium otophorum (Miquel) Koidzumi Fl. Symb. Orient.-Asiat. 40. 1930.
光蹄盖蕨 guang ti gai jue
Asplenium otophorum Miquel, Ann. Mus. Bot. Lugduno-Batavi 3: 175. 1867; Athyrium erythrocaulon Ching; A. rigescens Makino; A. sinovidalii Ching & Z. Y. Liu; A. vidalii Nakai var. chinense Koidzumi; A. violascens Diels; Diplazium otophorum (Miquel) C. Christensen.
Rhizomes short, ascending, apex densely clothed with brown or blackish brown, linear-lanceolate, fibriform-tipped scales. Fronds caespitose; fertile fronds 45-70(-85) cm; stipe blackish brown, upward slightly purplish red, (15-)25-35 cm, 2.5-3 mm in diam., base densely clothed with scales, upward glabrate; lamina 2-pinnate, narrowly ovate or deltoid-ovate, 25-35(-50) × 20-25 cm at middle, base not narrowed, apex abruptly narrowed; pinnae ca. 15 pairs, under abruptly narrowed tip ca. 7 pairs, basal pinnae opposite, upper pinnae alternate, subspreading, sessile or very shortly stalked, lanceolate; middle pinnae 10-12(-25) × 2.5-3.5 cm, base truncate, pinnate, apex long acuminate; pinnules 14-17 pairs, alternate, subspreading, sessile, lower pinnules subtriangular to oblong-lanceolate, 1-1.7 cm × 4-6 mm at middle, base inequilateral, truncate, parallel to costae, auriculate on acroscopic side, auricles triangular, cuneate on basiscopic side, subentire or with small tooth at acroscopic margin, apex acute; veins visible abaxially, inconspicuous adaxially, pinnate in pinnules, lateral veins 7 or 8 pairs, oblique, lower veins forked, basal acroscopic veins 2-pinnate or 3-forked. Lamina papery when dried, pale brown, glabrate on both surfaces; rachis and costae pale red-purple, glabrate abaxially, with subulate spines on adaxial side at base of costa or costule. Sori oblong or shortly linear, 3-5 pairs per pinnule, inframedial; indusia brownish, oblong or shortly linear, membranous, entire, persistent. Perispore surface without folds, with granular ornamentation. 2n = 160.
Evergreen broad-leaved forests, bamboo forests, wet areas; 400-1400(-3000) m. Anhui, Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Sichuan, Taiwan, Yunnan, Zhejiang [Japan, Korea].
Knapp (Ferns Fern Allies Taiwan, 439. 2011) noted that Athyrium otophorum is closely allied with A. delavayi, A. epirachis, and A. pubicostatum; and, although it is often distinguished by the glabrous abaxial surface of the rachis and costae, sessile pinnae, and auriculate pinnule bases, its diagnostic characters are not always definite.