腺毛马蓝Pteracanthus forrestii (Diels) C. Y. Wu
腺毛马蓝Pteracanthus forrestii (Diels) C. Y. Wu
14．腺毛马蓝（中国高等植物图鉴）图版20：1-6
Pteracanthus forrestii (Diels) C. Y. Wu, Index Fl. Yunnan.(云南种子植物名录)2: 1682. 1984. ——Strobilanthes forrestii Diels in Not. Bot. Gard. Edinb. 5: 162. 1912;中国高等植物图鉴4: 161. 1975; non est Gutzlaffia forrestii S. Moore in Journ. Bot. 63: 167. 1925; Bremek. in Verh. Ned. Akad. Wetensch. Afd. Naturk. Sect. 2. 41(1): 275. 1944; E. Hossain in Lauener, Not. Roy. Bot. Gard. Edinb. 38(3): 478. 1980; J. R. I. Wood in Edinb. Journ. Bot. 51(2): 208, 209, 238, 268. 1994. ——Strobilanthes panpienkaiensis Levl. Cat. Yunn. : 6. 1915; R. Ben. in Fedde Bull. Mus. Naturk. His. Naturk. 28: 98. 1922; Bremek. in Verh. Ned. Akad. Wetensch. Afd. Naturk. Sect. 2. 41(1): 275. 1944; J. R. I. Wood in Edinb. Journ. Bot. 51(2): 238. 1994.
灌木或草本，高达1米，植株遍生柔毛和腺毛，后毛脱落。叶草质，几无柄，卵形至卵状矩圆形，基部宽，有时近心形，稀楔形而狭，顶端略钝至略尖，长2-5厘米，宽1.2-3厘米，边有锯齿。花序穗状，长5-15厘米，或基部有分枝，每节具对生双花，节间长1-2.5厘米，近圆锥花序；花单生，无梗。苞片叶状，上部的全缘，卵状椭圆形，长1-3厘米；小苞片条形，狭矩圆形，等长或短于花萼裂片；花萼裂片5,条形，长8-12毫米，其中1片稍长，花萼连同苞片和小苞片密被腺状微柔毛；花冠蓝色或紫色或白色，长约3-3.5厘米，花冠管基部细狭，上部扩大钟状圆柱形，并弯曲，外面疏生微毛，里面有2列柔毛及背部疏生微毛，冠檐裂片5，几相等，长约3毫米；雄蕊4，2强，花丝基部有膜相连，药室平行；花粉粒超长球形（据冯国楣9024云南丽江；李国凤60057四川南川标本），具3孔沟及假沟，带形纹饰，带由环形凸起组成。花柱顶端稍扩大，子房顶端有微腺毛。蒴果长约1.2厘米，顶部有毛；种子4粒，深褐色，有微毛，基区白色，小。
产云南（大理、丽江、中甸）。生林下或草坡。在滇西北的亚高山针叶林下常可成大片生长，海拔几达3000米。选模式采自云南丽江。
J. R. I. Wood在Edinb. Journ. Bot. 51（2）：238. 1994.文中，选定G. Forrest 2760为本种选模式，并检验了Diels原引的2456, 4551, 4553四号标本，认为Forrest 2456实际上是Strobilanthes versicolor，原因有二，一是Diels在同一篇文中将其描述为Strobilanthes versicolor，二是该号标本叶也有柄，花萼相等。
《Flora of China》 Vol. 19 (2011)
Strobilanthes forrestii Diels Notes Roy. Bot. Gard. Edin-burgh. 5: 162. 1912.
腺毛马蓝 xian mao ma lan
Pteracanthus forrestii (Diels) H. P. Tsui; P. rotundifolius (Benoist) Bremekamp; Strobilanthes panpienkaiensis H. Léveillé; S. rotundifolia Benoist.
Herbs to 30-75 cm tall, isophyllous. Stems 4-angled, thinly pilose with gland-tipped trichomes, glabrescent, basally woody. Leaves subsessile; leaf blade ovate to rarely ovate-oblong, 2-5 × 1.2-3 cm, both surfaces thinly pilose, adaxially with cystoliths, secondary veins ca. 4 on each side of midvein, base rounded, cordate, or rarely cuneate, margin crenate to subentire, apex acute to obtuse. Inflorescences terminal or very rarely also from apical leaf axils, spikes, 5-15 cm; rachis pilose with gland-tipped trichomes; bracts leaflike, pilose with gland-tipped trichomes; basal bracts ovate-elliptic, 1-3 × 1-2 cm, margin obscurely crenate; apical bracts lanceolate, 7-10 × ca. 3 mm, margin entire; bracteoles linear to narrowly oblong, 6-11 × ca. 1 mm, pilose with gland-tipped trichomes. Flowers 1-2.5 cm apart on rachis. Calyx 0.8-1.2 cm, 5-lobed almost to base; lobes linear with one ca. 2 mm longer than others, pilose with gland-tipped trichomes. Corolla blue to purple, 3.2-4 cm, curved, outside with a few trichomes, inside glabrous except for trichomes retaining style; tube basally cylindric and ca. 2 mm wide for ca. 5 mm then bent to ca. 90° and widened to ca. 1.5 cm at mouth; lobes ovate, ca. 6 × 7 mm, apex obtuse. Stamens 4, included; shorter filament pair ca. 2 mm; longer filament pair ca. 8 mm, shortly pubescent; anther thecae oblong, ca. 2 × 0.8 mm; pollen type 3. Ovary gland-tipped puberulent at tip; style ca. 2.8 cm. Capsule oblong, ca. 1.2 cm, apically gland-tipped pilose, 4-seeded. Seeds dark brown, ovate in outline, ca. 3 mm, pubescent; areola white, small. Fl. Jul-Aug.
● Forests, grass slopes; ca. 3000 m. Yunnan.
Strobilanthes forrestii was misidentified as S. extensa (Pteracanthus extensus) by Handel-Mazzetti (Symb. Sin. 7: 895. 1936), C. Y. Wu (Index Fl. Yunnan. 2: 1682. 1984), and H. P. Tsui and C. C. Hu (FRPS 70: 136. 2002; Fl. Yunnan. 16: 696. 2006).Strobilanthes forrestii was misidentified as S. extensa (Pteracanthus extensus) by Handel-Mazzetti (Symb. Sin. 7: 895. 1936), C. Y. Wu (Index Fl. Yunnan. 2: 1682. 1984), and H. P. Tsui and C. C. Hu (FRPS 70: 136. 2002; Fl. Yunnan. 16: 696. 2006).
别名：味牛夕;白升麻;味牛膝;牛克膝;腺毛马兰;毛叶草;土牛夕;
科名：爵床科 Acanthaceae
属名：马蓝属 Pteracanthus
14．腺毛马蓝（中国高等植物图鉴）图版20：1-6
Pteracanthus forrestii (Diels) C. Y. Wu, Index Fl. Yunnan.(云南种子植物名录)2: 1682. 1984. ——Strobilanthes forrestii Diels in Not. Bot. Gard. Edinb. 5: 162. 1912;中国高等植物图鉴4: 161. 1975; non est Gutzlaffia forrestii S. Moore in Journ. Bot. 63: 167. 1925; Bremek. in Verh. Ned. Akad. Wetensch. Afd. Naturk. Sect. 2. 41(1): 275. 1944; E. Hossain in Lauener, Not. Roy. Bot. Gard. Edinb. 38(3): 478. 1980; J. R. I. Wood in Edinb. Journ. Bot. 51(2): 208, 209, 238, 268. 1994. ——Strobilanthes panpienkaiensis Levl. Cat. Yunn. : 6. 1915; R. Ben. in Fedde Bull. Mus. Naturk. His. Naturk. 28: 98. 1922; Bremek. in Verh. Ned. Akad. Wetensch. Afd. Naturk. Sect. 2. 41(1): 275. 1944; J. R. I. Wood in Edinb. Journ. Bot. 51(2): 238. 1994.
灌木或草本，高达1米，植株遍生柔毛和腺毛，后毛脱落。叶草质，几无柄，卵形至卵状矩圆形，基部宽，有时近心形，稀楔形而狭，顶端略钝至略尖，长2-5厘米，宽1.2-3厘米，边有锯齿。花序穗状，长5-15厘米，或基部有分枝，每节具对生双花，节间长1-2.5厘米，近圆锥花序；花单生，无梗。苞片叶状，上部的全缘，卵状椭圆形，长1-3厘米；小苞片条形，狭矩圆形，等长或短于花萼裂片；花萼裂片5,条形，长8-12毫米，其中1片稍长，花萼连同苞片和小苞片密被腺状微柔毛；花冠蓝色或紫色或白色，长约3-3.5厘米，花冠管基部细狭，上部扩大钟状圆柱形，并弯曲，外面疏生微毛，里面有2列柔毛及背部疏生微毛，冠檐裂片5，几相等，长约3毫米；雄蕊4，2强，花丝基部有膜相连，药室平行；花粉粒超长球形（据冯国楣9024云南丽江；李国凤60057四川南川标本），具3孔沟及假沟，带形纹饰，带由环形凸起组成。花柱顶端稍扩大，子房顶端有微腺毛。蒴果长约1.2厘米，顶部有毛；种子4粒，深褐色，有微毛，基区白色，小。
产云南（大理、丽江、中甸）。生林下或草坡。在滇西北的亚高山针叶林下常可成大片生长，海拔几达3000米。选模式采自云南丽江。
J. R. I. Wood在Edinb. Journ. Bot. 51（2）：238. 1994.文中，选定G. Forrest 2760为本种选模式，并检验了Diels原引的2456, 4551, 4553四号标本，认为Forrest 2456实际上是Strobilanthes versicolor，原因有二，一是Diels在同一篇文中将其描述为Strobilanthes versicolor，二是该号标本叶也有柄，花萼相等。
《Flora of China》 Vol. 19 (2011)
Strobilanthes forrestii Diels Notes Roy. Bot. Gard. Edin-burgh. 5: 162. 1912.
腺毛马蓝 xian mao ma lan
Pteracanthus forrestii (Diels) H. P. Tsui; P. rotundifolius (Benoist) Bremekamp; Strobilanthes panpienkaiensis H. Léveillé; S. rotundifolia Benoist.
Herbs to 30-75 cm tall, isophyllous. Stems 4-angled, thinly pilose with gland-tipped trichomes, glabrescent, basally woody. Leaves subsessile; leaf blade ovate to rarely ovate-oblong, 2-5 × 1.2-3 cm, both surfaces thinly pilose, adaxially with cystoliths, secondary veins ca. 4 on each side of midvein, base rounded, cordate, or rarely cuneate, margin crenate to subentire, apex acute to obtuse. Inflorescences terminal or very rarely also from apical leaf axils, spikes, 5-15 cm; rachis pilose with gland-tipped trichomes; bracts leaflike, pilose with gland-tipped trichomes; basal bracts ovate-elliptic, 1-3 × 1-2 cm, margin obscurely crenate; apical bracts lanceolate, 7-10 × ca. 3 mm, margin entire; bracteoles linear to narrowly oblong, 6-11 × ca. 1 mm, pilose with gland-tipped trichomes. Flowers 1-2.5 cm apart on rachis. Calyx 0.8-1.2 cm, 5-lobed almost to base; lobes linear with one ca. 2 mm longer than others, pilose with gland-tipped trichomes. Corolla blue to purple, 3.2-4 cm, curved, outside with a few trichomes, inside glabrous except for trichomes retaining style; tube basally cylindric and ca. 2 mm wide for ca. 5 mm then bent to ca. 90° and widened to ca. 1.5 cm at mouth; lobes ovate, ca. 6 × 7 mm, apex obtuse. Stamens 4, included; shorter filament pair ca. 2 mm; longer filament pair ca. 8 mm, shortly pubescent; anther thecae oblong, ca. 2 × 0.8 mm; pollen type 3. Ovary gland-tipped puberulent at tip; style ca. 2.8 cm. Capsule oblong, ca. 1.2 cm, apically gland-tipped pilose, 4-seeded. Seeds dark brown, ovate in outline, ca. 3 mm, pubescent; areola white, small. Fl. Jul-Aug.
● Forests, grass slopes; ca. 3000 m. Yunnan.
Strobilanthes forrestii was misidentified as S. extensa (Pteracanthus extensus) by Handel-Mazzetti (Symb. Sin. 7: 895. 1936), C. Y. Wu (Index Fl. Yunnan. 2: 1682. 1984), and H. P. Tsui and C. C. Hu (FRPS 70: 136. 2002; Fl. Yunnan. 16: 696. 2006).Strobilanthes forrestii was misidentified as S. extensa (Pteracanthus extensus) by Handel-Mazzetti (Symb. Sin. 7: 895. 1936), C. Y. Wu (Index Fl. Yunnan. 2: 1682. 1984), and H. P. Tsui and C. C. Hu (FRPS 70: 136. 2002; Fl. Yunnan. 16: 696. 2006).